Eimer similarly denies any capacity on the part of natural selection to
create species.
Pfeffer urges a very different objection. He says that if such a force as
natural selection existed it would transform species much more rapidly
than it does!
Now, in order that the above objections can carry any weight, one of two
sets of conditions must be fulfilled.
Either all organisms must be perfectly adapted to their environment, and
this environment must never change, or there must be inherent in each
species a kind of growth-force which impels the species to develop in
certain fixed directions. In either of these circumstances natural
selection will be an inhibitory force, for if the normal organism is
perfectly adapted to its environment, all variations from the type must
be unfavourable, and natural selection will weed out the individuals that
display them. No careful student of nature can maintain, either that all
animals are perfectly adapted to their environment, or that this never
changes. Hence those who deny that natural selection is a factor in the
making of species, assume the second set of conditions, that species
develop in certain fixed directions, being impelled either by internal or
external forces. How far these ideas are founded on fact we shall
endeavour to determine when speaking of variation. It must suffice at
present to say that even if any of these views of orthogenesis be
established, natural selection will have, so to speak, a casting vote, it
will decide which series of species developing along preordained lines
shall survive and which shall not survive.
Thus we reach by a different line of argument the conclusion we arrived
at in the last chapter: namely, there is no room for doubt that natural
selection is a factor in the making of species.
We must now pass on to the second class of objections, those which are
urged against the all-sufficiency of natural selection. So numerous are
these that it is not feasible to consider them all. A brief notice of the
more important ones should suffice to satisfy any unbiassed person;
firstly, that natural selection is an important factor in evolution;
secondly, that the position taken up by Wallace and his followers, that
natural selection, acting on minute variations, is the one and only
factor in organic evolution, is untenable.
Darwinism does not explain Variation
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