Thus, except in the very improbable case of a family of hybrids going off
and starting a colony by themselves, the effect of hybridism on the
evolution of species seems likely to have been _nil_. It is, however,
curious that three-quarter-bred animals have rarely, if ever, been
recorded in a state of nature, though a good many wild-bred hybrids are
on record.
This points to some unfitness for the struggle for existence even in a
fertile hybrid. It is necessary to emphasise the fact that wild hybrids
are always exceedingly rare as individuals, in spite of what has been
said as to the number of recorded crosses.
More hybrid unions have been noted among the duck family than anywhere
else in the animal kingdom. Nevertheless Finn never once saw a hybrid
duck for sale in the Calcutta market, although for seven years he was
constantly on the look-out for such forms; nor does Hume record any such
specimen in his _Game Birds and Wild Fowl of India_.
The hybrid which occurs most commonly as an individual is that between
the blackcock and capercailzie, which is recorded yearly on the
Continent; but it appears to be sterile, and so has no influence on the
species.
Wild hybrids between mammals are far rarer even than bird hybrids, the
only ones which seem to be on record being those between the Argali and
Urial above alluded to; those between the brown and blue hares and the
common and Arctic foxes.
A consideration of the phenomena of hybridism thus leads us to the
conclusion that, although many hybrids are fertile, the crossing of
distinct species has exercised little or no effect on the origin of
species. Even where allied species, like the pintail and the mallard
ducks, whose hybrid offspring is known to be fertile, inhabit the same
breeding area and occasionally interbreed in nature, such crossing does
not, for some reason or other, appear to affect the purity of the
species.
Very different, of course, is the effect of crossing a mutation within a
species with the parent form; the offspring are, as we shall see, likely
to resemble one or other of the parents; so that, if the mutation occur
frequently enough and be favourable to the species, the new form may in
course of time replace the old one.
CHAPTER V
INHERITANCE
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