Only to a certain extent can it be safely contended that the present route of a species is an indication of its earlier journeys, or that the direction of original dispersal is recapitulated in the present line of migration. Heredity, experience, and imitation would certainly tend to preserve and confirm the general direction; the shortest and easiest passage from food-base to food-base would become an hereditary route, unless circumstances arose which caused a change. Mr Cooke shows how there has probably been evolution of the route as well as of everything else concerned with a mutable animal. The fly-line across an arm of the sea may be lengthened if this lengthening means a corresponding advantage in reaching the desired haven. Thus the birds which now cross the Gulf of Mexico at its widest part, at one time probably coasted round the Gulf, as many do still, by the land-bridge of Mexico and Central America. The gradual straightening of this curve would shorten the journey both in time and distance, though lengthening the actual single flight across a portion of the sea. We can imagine a bird arriving in autumn at the mouth of the Mississippi, at first passing from Louisiana to Mexico, so as to save the time of travel through Texas. Generations later the shortening of the journey, through lengthening of the short cut, would lead the birds to Vera Cruz and later still to Yucatan. It may be questioned, what object could the birds have in risking an oversea voyage, away from chance of food and hope of rest, when the land-bridge remained open for them? Each individual or group of individuals which arrived at any particular place a little in advance of the migrating multitudes of its own species, or others which fed upon the same kind of food, would certainly gain advantage, and would be the most likely to develop strong flight and the power of endurance in its descendants; it would indeed be a winner in life's race. Great weight has been placed upon the use of land-bridges and the hereditary habit of crossing seas where these land-bridges once existed but have been submerged during the great geological changes in the earth's surface. Many have insisted that wherever migrants cross the sea they do so along submerged coast-lines or over submerged land-bridges, arguing that the gradual evolution which has made the advantageous adoption of a habit of migration possible was unable to eliminate the hereditary tendency to follow the exact route by which their ancestors passed from place to place. That there have been considerable alterations in coast-lines and in the general distribution of land and water since the time when birds began to be migratory is indeed probable, but unless crossing the sea means a distinct advantage it implies the retention of a habit which would not only be useless but might be a positive danger to the species.