The Movements and Habits of Climbing PlantsDarwin, Charles
Science
The Movements and Habits of Climbing Plants
Darwin, Charles
Climbing plants; Geotropism; Plants; Plants -- Irritability and movements
length, whilst the longest tendril was nearly double this length, namely
16 inches.
The gradations from the ordinary state of a flower-stalk, as represented
in the drawing (fig. 10), to that of a true tendril (fig. 9) are
complete. We have seen that the sub-peduncle (C), whilst still bearing
from thirty to forty flower-buds, sometimes becomes a little elongated
and partially assumes all the characters of the corresponding branch of a
true tendril. From this state we can trace every stage till we come to a
full-sized perfect tendril, bearing on the branch which corresponds with
the sub-peduncle one single flower-bud! Hence there can be no doubt that
the tendril is a modified flower-peduncle.
Another kind of gradation well deserves notice. Flower-tendrils (B, fig.
10) sometimes produce a few flower-buds. For instance, on a vine growing
against my house, there were thirteen and twenty-two flower-buds
respectively on two flower-tendrils, which still retained their
characteristic qualities of sensitiveness and spontaneous movement, but
in a somewhat lessened degree. On vines in hothouses, so many flowers
are occasionally produced on the flower-tendrils that a double bunch of
grapes is the result; and this is technically called by gardeners a
“cluster.” In this state the whole bunch of flowers presents scarcely
any resemblance to a tendril; and, judging from the facts already given,
it would probably possess little power of clasping a support, or of
spontaneous movement. Such flower-stalks closely resemble in structure
those borne by _Cissus_. This genus, belonging to the same family of the
Vitaceæ, produces well-developed tendrils and ordinary bunches of
flowers; but there are no gradations between the two states. If the
genus _Vitis_ had been unknown, the boldest believer in the modification
of species would never have surmised that the same individual plant, at
the same period of growth, would have yielded every possible gradation
between ordinary flower-stalks for the support of the flowers and fruit,
and tendrils used exclusively for climbing. But the vine clearly gives
us such a case; and it seems to me as striking and curious an instance of
transition as can well be conceived.
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