Another most important conclusion to be derived from the uprising of the
Amphibia is that at that time there was no race of animals living on the
land which had a chance against them. No race of land-living animals had
been evolved whose organization enabled them to compete with and overcome
these intruders from the sea in the struggle for existence. For this reason
that the whole land was their own, and no serious competition could arise
from their congeners, the fish, they took possession of it, and increased
mightily in size; losing more and more the habit of going into the water,
becoming more and more truly terrestrial animals. Henceforth, then, in
trying to find out the sequence of evolution, we must leave the land and
examine the nature of the animals living in the sea; the air-breathing
animals which lived on the land in the Upper Silurian and Devonian times
cannot have reached a stage of organization comparable with that of the
fishes, seeing how easily the amphibians became dominant.
We arrive, then, at the conclusion that the ancestors of the fishes must
have lived in the sea, and applying still the same principles that have
held good up to this time, the ancestors of the fishes must have arisen
from some member of the race predominant at the time when they first
appeared, and also the earliest fishes must have much more closely
resembled the ancestral form than those found in later times or at the
present day.
What, then, is the record of the rocks at the time of the first appearance
of fish-like forms? What kind of fishes were they, and what was the
predominant race at the time?
We have now reached the Upper Silurian and Lower Devonian times, and most
instructive and suggestive is the revelation of the rocks. Here, when the
first vertebrates appeared, the sea was peopled with corals, brachiopods,
early forms of cephalopods, and other invertebrates; but, above all, with
the great tribe of trilobites (Fig. 6) and their successors. From the
trilobites arose, as evidenced by their larval form, the king-crab group,
called the Xiphosura (Fig. 5). Closely connected with them, and forming
intermediate stages between trilobites and king-crabs, numerous forms have
been discovered, known as Belinurus, Prestwichia, Hemiaspis, Bunodes, etc.
(Fig. 5 and Fig. 12). From them also arose the most striking group {25}of
animals which existed at this period--the giant sea-scorpions, or
Gigantostraca. This group was closely associated with the king-crabs, and
the two groups together are classified under the title Merostomata.
[Illustration: FIG. 5 (from H. WOODWARD).--1. _Limulus polyphemus_ (dorsal
aspect). 2. _Limulus,_ young, in trilobite stage. 3. _Prestwichia
rotundata._ 4. _Prestwichia Birtwelli._ 5. _Hemiaspis limuloides._ 6.
_Pseudoniscus aculeatus._]
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