The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
entirely rejects Quenstedt's opinions with regard to their upright
position. He makes the following remarks: 'The animal walked on the
free fore-toes and bore the wings like a bat, though with the body not
in an upright position like a bird, but four-footed. The hind-foot is
much too small for such an upright position, and the fore-foot much too
strongly developed. I therefore believe that the Pterodactyle could
much better have walked four-footed than a bat, because it possessed so
much better developed fore-feet.' In the length of the tibia Burmeister
sees no reason for the upright position, but, as he says, only a means
for the wide expansion of the flying-membrane;--and an endeavour in
walking on four feet to bring the leg into the necessary harmony with
the arm, which is so much elongated with the flat-hand.
Hermann von Meyer[L].
[Footnote L: _Fauna der Vorwelt. Reptilien aus dem Lithographischen
schiefer._ Frankfurt am Main. 1859. pp. 15-23.]
The skull of the Pterodactyle can only be compared with those of
birds and lizards. The form is essentially Avian, and the sutures are
indistinct or obliterated as in birds, while in reptiles they are
persistent The temporal bone enters into the formation of the reservoir
for the brain, which is eminently characteristic of birds and quite
different from anything found in lizards. The snout resembles a bird in
being chiefly formed by the intermaxillary bone, which bounds the front
of the anterior nares; and, as in birds, the bone extends backward
between the eye-cavities to the frontal bone. The corresponding
intermaxillary ridge of the Monitor is of less extent.
The frontal-bone forms the highest part of the skull, and is similar
to that of birds. The principal frontal is double, and forms the upper
and hind part of the cavity for the eye, and covered the greater part
of the large brain, composed of two hemispheres, in which Oken long
ago saw a similarity to the higher animals. The arched form of the
back part of the skull is bird-like. The double parietal adjoins the
principal frontal, and is conditioned like the parietal in birds. The
supra-occipital is single as in birds, expanded, and forms the part of
the skull which extends furthest back. From the form of the back part
of the skull it may be concluded that the foramen magnum was situated
as in birds, and that the head and neck were moved as in birds, and not
as in reptiles and mammals.
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