The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
In length the metacarpus resembles that of the Ruminants, in which
however it consists of but one bone; while in Pterodactyles there is
a separate bone for each of the four fingers; they are closely united
together without being blended. In some Pterodactyles the metacarpals
of the short fingers are as fine as hairs, so that it is impossible
that they should have articular facets on the carpus. In Ornithopterus
the metacarpus has some resemblance with that of the bird, but the
articulation with the phalanges of the finger for flight is stiff. In
Pterodactylus and Rhamphorhynchus there is a free articulation.
Burmeister remarks that the chief articulation of the wing in bats is
with the carpus, while in Pterodactyle the articulation is with the end
of the metacarpus.
_The hand._
Von Meyer finds four fingers. It was formerly supposed that the order
of the phalanges was 2, 3, 4, 4, but in the fly-finger this is not the
case, Ornithopterus having but two. The number of joints in the other
fingers is quite as irregular.
In Pt. longicollum the thumb consists of but one joint.
_The Ilium_
is more mammalian and avian than reptilian.
_Pubis._
The pubis appears to have been excluded from the glenoid cavity, as
in Crocodiles. It is more mammal-like than bird-like, and is to be
compared with the marsupial bones.
_The femur._
In certain Pterodactyles the proximal condyle of the femur resembles
birds; but in other Pterodactyles the bone is more mammal-like in its
straightness, and development of the upper condyle, and in the presence
of a trochanter.
_The tibia and fibula_
may be compared, from their great length, with birds and flying
vertebrate animals.
The fibula is style-shaped, like that of a bird, the lower part being
wanting; while in bats the upper part is wanting.
_The tarsus,_
of two rows, is best compared with that of reptiles. The number of
constituent bones has not been definitely determined.
_The metatarsus_
shows a certain return from the bird type to that of reptiles.
_Foot._
Von Meyer never finds more than four toes, and sometimes a stump of a
fifth. As a whole, the foot is Saurian-like. It differs from lizards in
the number of toes, and approximates to Crocodiles. In Pterodactylus
longirostris the formula of the toes is 2, 3, 4, 5, with a stump of two
joints;--like lizards, if we abstract the outer toe; and like birds
with four toes; but they are liable to variations.
In Pterodactylus scolopaciceps and P. Kochi the formula is 2, 3, 3,
4 joints. In Winkler's specimen of P. Kochi there is also a stump of
three joints.
In Pterodactylus micronyx the formula is 2, 3, 3, 3, and a stump of two
joints. In P. longicollum the number appears to be different from all
the foregoing.
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