The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
Pl. 4.
Case. Comp. Tablet. Specimen.
=J= _a_ 6 1-46
7 1- 3
8 1
There are among the fossils of the Cambridge Greensand at least two
well-marked _types_ of Pterodactyle humerus, readily recognised by the
forms of the proximal and of the distal ends, and by the positions of
the pneumatic foramina. In the group having the ulnar ridge developed
the pneumatic foramen is on the posterior aspect of the bone[O] under
the ulnar ridge, as in birds; but in some of the small Pterodactyles
the foramen is on the anterior surface, and on its radial side. This
latter kind of humerus has the distal end more or less divided into
three convex surfaces, while the radial crest is enormously developed
and terminates in a smooth oblong flattened surface nearly as large
as the proximal articular surface, and looking anteriorly. The distal
articular surfaces are not as in birds parallel to that of the proximal
end, though they agree with those of birds in being at right angles
to the radial crest; this ridge in Pterodactyles being directed much
further outward and backward than in birds.
[Footnote O: Professor Owen states (p. 16, 3d Supt.) that the foramen
is palmar. Fig. 15. T. III. 2d Supt. shows it to be anconal.]
The largest forms of Pterodactyle all have the distal articular surface
flatter, and the proximal articulation less bent back so as to look
more upwards. No specimen of this kind of humerus has occurred with the
radial crest preserved; but it is apparently carried farther down the
shaft and not so far forward as in the other group. This latter kind
of bone is shown by Prof. Owen in T. III. figs. 1, 2, 3rd Sup. Cret.
Reptiles; the former kind has been illustrated in figure 5 of the same
plate.
Some of the most gigantic Pterodactyles appear to have had the
limb-bones as solid as those of crocodiles, and unpermeated by air;
and there is no evidence that the high Avian characteristics of most
of these Greensand fossils also pertained to all the previously known
types from the lower secondary rocks.
The osteological series comprises 46 specimens. No. 30 is a nearly
perfect right humerus. Nos. 1, 2, 5, 6, 7, 8, 9, 11, 18, 22, 23, 25, 39
are examples of the proximal ends of left humeri. Nos. 3, 4, 10, 12,
13, 14, 15, 16, 17, 19, 24, 26, 27, 28, 38, 40, 41 are examples of the
proximal ends of right humeri. Nos. 20, 21, 32, 33, 34, 35, 37, 42, 44,
45, are examples of the distal ends of left humeri. Nos. 29, 31, 36, 43
and 46 are distal ends of right humeri.
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