In this assumption we postulate for biology the
distinctive mathematical concept of infinite divisibility.
[29] See Appendix, p. 350.
The difference from the mode, or mean, with respect to a definite
character in a fully grown organism may be due to the direct action of
the environment, in the sense in which we have regarded the environment
as influencing the organism; or it may be due to the changes in the
organism resulting from the increased or decreased use of some of its
parts. The conditions with regard to nutrition, for instance, will not
be the same for all the individuals composing a cluster of mussels
growing on the sea-bottom. Those in the interior of the cluster do not
receive so abundant a supply of sea-water as those on the outside of
the cluster; and since the amount of food received by any individual
depends on the quantity of water streaming over it in unit time, we
shall find that the internally situated individuals will be stunted
or dwarfed, while those on the outside will be well grown. Such
variations are acquired ones, but even when we allow for them, even
if we take care that all the organisms studied live under conditions
which are as nearly uniform as possible, there will still be some
degree of variability. We cannot be sure that this absolute uniformity
ever exists; and the notion of the environment of an organism may be
extended so as to include the medium in which embryonic development
took place, and even the parental body which formed the environment
for the germ-cells from which embryonic development began. But it is
probably the case that even with an uniform environment, or with one
in which the differences were insignificant, variability would still
exist. The variations that might be observed in such a case would
belong to two kinds--“fluctuating variations,” and “mutations.”
[Illustration: FIG. 21.]
Whether the variations observed in a population of organisms are
fluctuations or mutations can only be determined by experiment.
Let us suppose that we are dealing with a human population, and
that the variation studied is that of stature. Let the men with
statures considerably over the mean value marry the women who are
correspondingly tall, then it will be found that the children from
these unions will, when grown up, exhibit a stature which is greater
than that of the whole population, but not so great as that of their
parents--that is, regression towards the mean of the whole population
takes place.
This is shown in the above diagram, where the lines above and below
the mean one indicate the proportion (relative to the value or
frequency of the mean) of people of each grade of stature. The latter
is proportional to the distance from the mean measured along the
vertical line, distances below this line indicating statures below the
mean, and _vice versa_.
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