It is, however, not quite accurate to say that the organisation
persists unchanged from generation to generation. The offspring is
similar to the parent--that is, the organisation has been transmitted
unchanged. But the offspring also differs just a little from the
parent--that is to say, the organisation is modified by each
transmission. In these two statements we formulate in the simplest
manner the law of organic variability. Organisms may obviously be
arranged in categories in such a way that the individuals in any one
category resemble each other more closely than they resemble the
individuals belonging to another category. We may, by experimental
breeding, produce an assemblage of organisms all of which have had
a common ancestor, or a pair of ancestors. Now the individuals
composing such an assemblage would exhibit a close resemblance
to each other, such a resemblance as our categories of naturally
occurring organisms are seen to exhibit. We should also find that the
individuals of our naturally occurring assemblage would be able to
interbreed among themselves, just as in the case of the experimentally
produced population. It may be concluded, then, that the naturally
occurring population is also the product of a pair of ancestors. This
inter-fertility, as well as the close morphological resemblance of the
individuals, are the facts on which the hypothesis of the common origin
and unity of the assemblage, or species, is formed.
The morphological resemblance between the individuals, either in
the natural or the artificial populations, is not absolute. If we
take any single character capable of measurement we shall find that
it is variable from organism to organism. This important concept of
organic variability may be made more clear by a concrete example.
Examination of a large number of cockle shells taken from the same
restricted part of the sea-shore, and therefore belonging presumably
to the same race, will show that the number of the radiating ridges
on the shell varies from 19 to 27, and that the ratio of the length
to the depth of the shell also varies from 1 : 0.59 to 1 : 0.85. In
the former case the most common number of ridges is 23, and in the
latter case the most common ratio of length to depth is 1 : 0.71.
These are the characteristic or modal values of the morphological
characters in question, and the other or less commonly occurring
values are distributed symmetrically on either side of the mean or
modal value, forming “frequency distributions.”[29] The value of
the first character changes by unity in any distribution: obviously
there cannot be a fraction of a ridge; and this kind of variation is
called “discontinuous.” The value of the second character may change
imperceptibly, and it is therefore called “continuous,” a term which
is not strictly accurate, since in applying it we assume that the
numerical difference between two variates may be less than any finite
number, however small.
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