All this is merely a conceptual physical model of a process about which
we know nothing at all. It is meant to illustrate the view that the
organisation of a plant or animal is not something absolutely fixed and
invariable. The organism in respect of each recognisable and measurable
character oscillates about a point of stability, that is to say
exhibits fluctuating variations about the mean value of this character.
If the stability of the organisation is upset, so that it oscillates,
or fluctuates about a new centre, that is, if the variations deviate
in either direction from a new “type” or mean, a mutation has been
established. A mutation is not, therefore, necessarily a large
departure from “normality.” It is not necessarily a “discontinuous
variation,” nor a “sport” nor a “freak.” It is essentially a shifting
of the mean position about which the variations exhibited by the
organism fluctuate.
Such a mutation will, in general, involve the creation of an
“elementary species.” We have considered only one character, say
stature, in the above discussion, but it generally happens that the
assumption of a new centre of stability involves _all_ the characters
of the mutating organism. An elementary species therefore differs a
little in respect of all its characters from the species from which it
arose, or from the other elementary species near which it is situated.
This is what we do usually find in the cases of the “races,” or “local
varieties,” of any one common species of plant or animal. That we
do not recognise that most, or perhaps all, of the species known to
systematic biology are really composed of such local races is merely
because such results involve an amount of close investigation such as
has not generally been possible except in the few cases studied with
the object of proving such variability; or in the case of those species
which are studied with great attention to detail because of their
economic importance. Thus the herrings of North European seas can be
divided into such races, and it is possible for a person possessing
great familiarity with these fishes to identify the various races or
elementary species--that is, to name the locality from which the fish
were taken--by considering the characteristics in respect of which the
herrings of one part of the sea differ from those of other parts.
Public-domain text, read in full here on John Shaqi.
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