A parallel chronological sequence would also be suggested by the
results of embryology. This branch of biology shows us that
all animals pass through a series of stages in their individual
development, or ontogeny. The earlier stages represent a simple type of
structure, usually a hollow ball of cells, but as development proceeds,
the structure of the embryo becomes more and more complex. The process
of development is continuous in many animals, but in others (perhaps in
most) larval stages appear, that is, development is interrupted, and
the animal may lead for a time an independent existence similar to that
of the fully developed form. Often these larval stages suggest types
of structure lower than that of the fully developed animal into which
they transform. Even if larval stages may not appear in the ontogeny,
it is very often the case that the developing embryo exhibits traces,
or at least reminiscences, of the types of morphology characteristic
of the animals which are lower or less complex than itself; thus the
piscine gills appear during the development of the tailed Amphibian,
and even in that of the Mammal, and then vanish, or are converted into
organs of another kind. The individual thus passes through a series
of developmental stages of increasing complexity: it repeats, in its
ontogeny, the palæontological sequence in a distorted and abbreviated
form.
It is true that the evidence afforded by palæontology is very meagre.
The preservation of the remains of organisms in the stratified rocks is
a very haphazard process, and it depends for its success on a series
of conditions that are not always present. As the surface of the earth
becomes better known, our knowledge of the life of the past will become
fuller, but there can be little doubt that whole series of organisms
must have existed in the past, and that no recognisable traces of
these are known to us. There is also no doubt that the sequences
indicated by palæontology are very incomplete: they are obscured and
shortened by many conditions. The earlier embryologists entertained
hopes that the study of embryology would reveal the direction of the
evolutionary process in many groups of animals: if the organism repeats
in its ontogeny the series of stages through which it passed in its
phylogenetic development, then a close study of the embryological
process ought to disclose these stages. Although these hopes have
not been realised, there is yet sufficient truth in the doctrine of
recapitulation to enable us to state that there is a rough parallelism
between the palæontological and embryological sequences.
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