bodily segment bears a pair of jointed appendages, and also contains
a separate nerve-centre. The whole series of ganglia are connected
together by means of a nerve-cord, and the nervous system lies ventral
to the alimentary canal. The Vertebrata are also segmented animals,
but the segmentation is not apparent externally. The skeleton is an
internal one, and is built up round an axial rod or notochord. The
nervous system is situated dorsally to the alimentary canal. There are
two pairs of limbs.
Thus we set up an essential or schematic structure characteristic
of each phylum. These schemata have no real existence: they are
morphological types from which the actual bodily structure of the
animals in each phylum may be deduced. They represent the minimum of
parts which must be present in order that an animal may be placed in
the phylum to which we assume that it may belong. But these anatomical
parts need not actually be present in the fully developed organism:
thus there are Crustacea in which the body is not segmented, and in
which neither calcareous exo-skeleton nor jointed appendages are
present; and there are Vertebrata in which the limbs may be absent.
But in such cases we require evidence that the essential anatomical
characters which are absent in the fully developed animal have appeared
at some stage in its ontogeny, and this evidence is usually available.
Or if embryological evidence cannot be obtained, we require proof
that the animal can be traced backwards in time, by means of other
characters, to some form in which the missing structures reappear.
The schemata are thus the generalised or conceptual morphology of the
phyla. They are not the morphology of an individual organism, but they
include the morphology of the race.
Public-domain text, read in full here on John Shaqi.
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