There is an essential structure which we endeavour to assign to all
the animals of each phylum, and which is different from the structure
of the animals belonging to all other phyla. The Protozoa, which for
the present we regard as animals, are organisms the bodies of which
consist of single cells. These cells may become aggregated into
colonies, but they may as well exist apart from each other. They may
be enclosed in limy, siliceous, or cellulose skeletons or shells, or
they may possess limy or siliceous spicules in their tissues--these
parts are non-essential, and the schematic Protozoan is a cell
containing a single nucleus, and capable of independent existence.
The Porifera, and all the other phyla, include organisms the bodies of
which are made up of aggregates of cells. In the Porifera the cells,
which are specially modified in structure, are arranged to form the
internal walls of a “sponge-work” the cavities of which open to the
outside by series of pores through which water is circulated. The
bodies of the Cœlenterates are typically sacs formed by a double wall
of cells--endoderm and ectoderm. This sac opens to the exterior by a
single opening, or mouth, surrounded by a circlet of tentacles, and
its cavity is the only one contained in the body of the animal. The
Platyhelminth worms are animals the bodies of which are also composed
of ectodermal and endodermal tissues, between which is intercalated
another mesodermal tissue. They have a single digestive sac or
alimentary canal opening to the exterior by means of a mouth only; and
they all possess a complex, hermaphrodite, reproductive apparatus. In
all the other phyla there are also three principal layers or kinds of
tissue, but in addition to the cavity of the alimentary canal there
is also a body cavity, or cœlom, which is contained in the mesodermal
tissues. The Echinoderms are such cœlomate animals, but the alimentary
canal now opens to the exterior by means of both mouth and anus;
there are separate systems of vessels through which water and blood
circulate; the blood-vascular system of vessels is closed to the
exterior, the water-vascular system being open; and the integument is
armed by means of calcareous spines or plates. The Annelids are animals
with cylindrically shaped bodies, segmented so as to form numerous
joints. Each segment bears spines or hairs or appendages of some sort,
and also contains a separate nerve-centre. The alimentary canal opens
externally by a mouth and anus, and there is a spacious body cavity.
The Molluscs are unsegmented animals. The dorsal part of their bodies
contains the viscera, and is protected by a shell; while the ventral
part is modified for the purpose of locomotion. A fold of integument
hangs down all round the body and encloses a cavity in which the gills
are contained. The Arthropods are segmented animals. The body is armed
by a calcareous carapace or shell which forms the exo-skeleton. Each
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