Let us consider further a man walking along the ties, or sleepers, of
a railway track. The ties are at variable distances apart, so that
the steps of the walker must vary in length, being sometimes closer
together, sometimes further apart. The _mean_ step has a definite
length and requires the expenditure of a certain amount of energy, and
the condition that the man takes sometimes a long step and sometimes
a short one does not require that the energy expended on the steps
should be more than if every one of them were of the mean length, for
the additional energy that is required for the long steps is saved from
the short ones. That which operates here is the power of regulation
exercised by the walker regarded as a mechanism. There is no purely
inorganic process precisely similar to this. It might be thought that
the governor of a steam engine did very much the same thing, admitting
more steam into the cylinder when the load on the engine increases,
and _vice versa_. But the governor is a mechanism _designed_ to
compensate for variations _that are given in advance_. In the case of
the man walking on the railway track, entelechy operates by suspending
energetic happening (the muscular contractions of the short steps) when
necessary, and allowing it to proceed when necessary. Entelechy itself,
whatever it may be, need not be affected by these regulations.
The organism is therefore an aggregation of chemical substances
arranged in a typical manner. These substances possess energy in the
potential form, capable of undergoing transformation so that they may
give rise to other chemical substances--secretions, for instance--or
to energy in the kinetic form, that is, the movements of muscles. In
the resting organism these transformations do not take place: the
energy remains potential, so that chemical happening is suspended. In
the unfertilised ovum, for instance, nothing happens although all the
potentialities of segmentation are contained in the cell. If reactions
did occur in consequence of the chemical potentials contained in the
substances of the cells, the progress of these would be such as to lead
to the formation of substances in which potential energy was minimal,
and in which the original energy of the cell would be represented by
the un-co-ordinated kinetic energy of the molecules resulting from the
breakdown of the substances undergoing the chemical changes. This is
not what happens in the differentiation of the ovum: the developing
cell forms new substances from those of its inorganic medium similar
to the substances of which it is already composed, and then these
substances become arranged to produce the specific form of the organism
into which the ovum is about to develop.
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