The physiology of digestion considered with relation to the principles of dieteticsCombe, Andrew
Science
The physiology of digestion considered with relation to the principles of dietetics
Combe, Andrew
Diet; Digestion
Something more, however, than the mere action of the teeth and jaws is
required to prepare the morsel for being swallowed. If we take a bit of
dry biscuit or mealy potato into the mouth, and attempt to masticate
it, we encounter at first no small difficulty from the stiffness and
resistance of the dry mass, and feel instinctively that it would be
in vain to attempt to swallow it, until moistened either by continued
mastication or by the admixture of fluid from without. In ordinary
states of the system, accordingly, a fluid called _saliva_ or
_spittle_ is copiously secreted and poured into the mouth for
this very purpose; and the process by which its due admixture with the
contents of the mouth is accomplished is called the _insalivation_
of the food.
To provide this necessary fluid, and to connect its supply directly
with the process of mastication to which it is subservient,
several glands for its secretion have been placed in the immediate
neighbourhood of the mouth and jaws, in such a way that the latter
cannot be opened and shut without affording them a stimulus, and still
farther increasing the secretion which the presence of the morsel is
itself sufficient to begin. From this arrangement it follows, that
the more perfectly mastication is performed, the more thoroughly does
the morsel become impregnated with the salivary fluid, and the better
fitted is it rendered for subsequent deglutition and digestion.
The apparatus of mastication varies according to the kind of food on
which the animal is destined to live; but in the higher orders of
animals, it consists essentially of the parts already mentioned. In
some animals, however, which live on soft gelatinous food--as the
whale--no teeth are to be found, because their peculiar power is not
required. In others--as the granivorous or grain-eating birds--the
grinding or _triturating_ process is effected not in the mouth but
in the gizzard, where the food (mixed with gravel, which the animal
is instinctively impelled to swallow for the purpose) is effectually
bruised and softened down by the strong muscles which constitute the
greater part of its substance. In these instances, the gravel is the
grinding instrument, and without its presence digestion cannot be
carried on any more than it could in man without the agency of teeth.
The degree of mastication required varies also according to the mode
of life of the animal, and the digestibility of its food. Animal food,
for example, being easy of digestion, requires less mastication than
vegetable food, which is more difficult. This is so much the case, that
most animals which live on fresh vegetable matter spend half their
waking hours in _ruminating_ or re-masticating the food, which
they have already cropped and stored up for the purpose in one of their
four stomachs. To this necessary act in them, Providence seems to have
attached a high degree of gratification, for the very purpose of
insuring its regular performance.
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