Botany; Geotropism; Plants -- Irritability and movements
In the several foregoing cases one of the cotyledons is delayed in its
development, or reduced in size, or rendered rudimentary, or quite
aborted; but in other cases both cotyledons are represented by mere
rudiments. With Opuntia basilaris this is not the case, for both
cotyledons are thick and large, and the hypocotyl shows at first no
signs of enlargement; but afterwards, when the cotyledons have withered
and disarticulated themselves, it becomes thickened, and from its
tapering form, together with its smooth, tough, brown skin, appears,
when ultimately drawn down to some depth into the soil, like a root. On
the other
hand, with several other Cacteæ, the hypocotyl is from the first much
enlarged, and both cotyledons are almost or quite rudimentary. Thus
with Cereus Landbeckii two little triangular projections, representing
the cotyledons, are narrower than the hypocotyl, which is pear-shaped,
with the point downwards. In Rhipsalis cassytha the cotyledons are
represented by mere points on the enlarged hypocotyl. In Echinocactus
viridescens the hypocotyl is globular, with two little prominences on
its summit. In Pilocereus Houlletii the hypocotyl, much swollen in the
upper part, is merely notched on the summit; and each side of the notch
evidently represents a cotyledon. Stapelia sarpedon, a member of the
very distinct family of the Asclepiadeae, is fleshy like a cactus; and
here again the upper part of the flattened hypocotyl is much thickened
and bears two minute cotyledons, which, measured internally, were only
.15 inch in length, and in breadth not equal to one-fourth of the
diameter of the hypocotyl in its narrow axis; yet these minute
cotyledons are probably not quite useless, for when the hypocotyl
breaks through the ground in the form of an arch, they are closed or
pressed against one another, and thus protect the plumule. They
afterwards open.
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