The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
While then it is certain, _a priori_, that there cannot be growth in the
absence of such substances as those of which an organism consists; and
while it is equally certain that the amount of growth must primarily be
governed by the supply of these substances; it is not less certain that
extra supply will not produce extra growth, beyond a point very soon
reached. Deduction shows to be necessary, as induction makes familiar, the
truths that the value of food for purposes of growth depends not on the
quantity of the various organizable materials it contains, but on the
quantity of the material most needed; that given a right proportion of
materials, the pre-existing structure of the organism limits their
availability; and that the higher the structure, the sooner is this limit
reached.
§ 46. But why should the growth of every organism be finally arrested?
Though the rate of increase may, in each case, be necessarily restricted
within a narrow range of variation--though the increment that is possible
in a given time, cannot exceed a certain amount; yet why should the
increments decrease and finally become insensible? Why should not all
organisms, when supplied with sufficient materials, continue to grow as
long as they live? To find an answer to this question we must revert to the
nature and functions of organic matter.
In the first three chapters of Part I, it was shown that plants and animals
mainly consist of substances in states of unstable equilibrium--substances
which have been raised to this unstable equilibrium by the expenditure of
the forces we know as solar radiations, and which give out these forces in
other forms on falling into states of stable equilibrium. Leaving out the
water, which serves as a vehicle for these materials and a medium for their
changes; and excluding those mineral matters that play either passive or
subsidiary parts; organisms are built up of compounds which are stores of
force. Thus complex colloids and crystalloids which, as united together,
form organized bodies, are the same colloids and crystalloids which give
out, on their decomposition, the forces expended by organized bodies. Thus
these nitrogenous and carbonaceous substances, being at once the materials
for organic growth and the sources of organic energy, it results that as
much of them as is used up for the genesis of energy is taken away from the
means of growth, and as much as is economized by diminishing the genesis of
energy, is available for growth. Given that limited quantity of nutritive
matter which the pre-existing structure of an organism enables it to
absorb; and it is a necessary corollary from the persistence of force, that
the matter accumulated as growth cannot exceed that surplus which remains
undecomposed after the production of the required amounts of sensible and
insensible motion. This, which would be rigorously true under all
conditions if exactly the same substances were used in exactly the same
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