The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
proportions for the production of force and for the formation of tissue,
requires, however, to be taken with the qualification that some of the
force-evolving substances are not constituents of tissue; and that thus
there may be a genesis of force which is not at the expense of potential
growth. But since organisms (or at least animal organisms, with which we
are here chiefly concerned) have a certain power of selective absorption,
which, partially in an individual and more completely in a race, adapts the
proportions of the substances absorbed to the needs of the system; then if
a certain habitual expenditure of force leads to a certain habitual
absorption of force-evolving matters that are not available for growth; and
if, were there less need for such matters, the ability to absorb matters
available for growth would be increased to an equivalent extent; it follows
that the antagonism described does, in the long run, hold even without this
qualification. Hence, growth is substantially equivalent to the absorbed
nutriment, minus the nutriment used up in action.
This, however, is no answer to the question--why has individual growth a
limit?--why do the increments of growth bear decreasing ratios to the mass
and finally come to an end? The question is involved. There are more causes
than one why the excess of absorbed nutriment over expended nutriment must,
other things equal, become less as the size of the animal becomes greater.
In similarly-shaped bodies the masses, and therefore the weights, vary as
the cubes of the dimensions; whereas the powers of bearing the stresses
imposed by the weights vary as the squares of the dimensions. Suppose a
creature which a year ago was one foot high, has now become two feet high,
while it is unchanged in proportions and structure; what are the necessary
concomitant changes? It is eight times as heavy; that is to say, it has to
resist eight times the strain which gravitation puts upon certain of its
parts; and when there occurs sudden arrest of motion or sudden genesis of
motion, eight times the strain is put upon the muscles employed. Meanwhile
the muscles and bones have severally increased their abilities to bear
strains in proportion to the areas of their transverse sections, and hence
have severally only four times the tenacity they had. This relative
decrease in the power of bearing stress does not imply a relative decrease
in the power of generating energy and moving the body; for in the case
supposed the muscles have not only increased four times in their transverse
sections but have become twice as long, and will therefore generate an
amount of energy proportionate to their bulk. The implication is simply
that each muscle has only half the power to withstand those shocks and
strains which the creature's movements entail; and that consequently the
creature must be either less able to bear these, or must have muscles and
bones having relatively greater transverse dimensions: the result being
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