The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
§ 74b. One of the complexities within complexities was disclosed when it
was found that the protoplasm itself has a complicated structure. Different
observers have described it as constituted by a network or reticulum, a
sponge-work, a foam-work. Of these the first may be rejected; since it
implies a structure lying in one plane. If we accept the second we have to
conceive the threads of protoplasm, corresponding to the fibres of the
sponge, as leaving interstices filled either with liquid or solid. They
cannot be filled with a continuous solid, since all motion of the
protoplasm would be negatived; and that their content is not liquid seems
shown by the fact that its parts move about under the form of granules or
microsomes. But the conception of moving granules implies the conception of
immersion in a liquid or semi-liquid substance in which they move--not a
sponge-work of threads but a foam-work, consisting everywhere of septa
interposed among the granules. This is the hypothesis which sundry
microscopists espouse, and which seems mechanically the most feasible: the
only one which consists with the "streaming" of protoplasm. Ordinarily the
name protoplasm is applied to the aggregate mass--the semi-liquid, hyaline
substance and the granules or microsomes it contains.
What these granules or microsomes are--whether, as some have contended,
they are the essential living elements of the protoplasm, or whether, as is
otherwise held, they are nutritive particles, is at present undecided. But
the fact, alleged by sundry observers, that the microsomes often form rows,
held together by intervening substance, seems to imply that these minute
bodies are not inert. Leaving aside unsettled questions, however, one fact
of significance is manifest--an immense multiplication of surfaces over
which inter-action may take place. Anyone who drops into dilute sulphuric
acid a small nail and then drops a pinch of iron filings, will be shown, by
the rapid disappearance of the last and the long continuance of the first,
how greatly the increasing of surfaces by multiplication of fragments
facilitates change. The effect of subdivision in producing a large area in
a small space, is shown in the lungs, where the air-cells on the sides of
which the blood-vessels ramify, are less than 1/100th of an inch in
diameter, while they number 700,000,000. In the composition of every tissue
we see the same principle. The living part, or protoplasm, is divided into
innumerable protoplasts, among which are distributed the materials and
agencies producing changes. And now we find this principle carried still
deeper in the structure of the protoplasm itself. Each microscopic portion
of it is minutely divided in such ways that its threads or septa have
multitudinous contacts with those included portions of matter which take
part in its activities.
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