The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
Concerning the protoplasm contained in each cell, named by some cytoplasm,
it remains to say that it always includes a small body called the
centrosome, which appears to have a directive function. Usually the
centrosome lies outside the nucleus, but is alleged to be sometimes within
it. During what is called the "resting stage," or what might more properly
be called the growing stage (for clearly the occasional divisions imply
that in the intervals between them there has been increase) the centrosome
remains quiescent, save in the respect that it exercises some coercive
influence on the protoplasm around. This results in the radially-arranged
lines constituting an "aster." What is the nature of the coercion exercised
by the centrosome--a body hardly distinguishable in size from the
microsomes or granules of protoplasm around--is not known. It can scarcely
be a repelling force; since, in a substance of liquid or semi-liquid kind,
this could not produce approximately straight lines. That it is an
attractive force seems more probable; and the nature of the attraction
would be comprehensible did the centrosome augment in bulk with rapidity.
For if integration were in progress, the drawing in of materials might well
produce converging lines. But this seems scarcely a tenable interpretation;
since, during the so-called "resting stage," this star-like structure
exists--exists, that is, while no active growth of the centrosome is going
on.
Respecting this small body we have further to note that, like the cell as a
whole, it multiplies by fission, and that the bisection of it terminates
the resting or growing stage and initiates those complicated processes by
which two cells are produced out of one: the first step following the
fission being the movement of the halves, with their respective completed
asters, to the opposite sides of the nucleus.
§ 74c. With the hypothesis, now general, that the nucleus or kernel of a
cell is its essential part, there has not unnaturally grown up the dogma
that it is always present; but there is reason to think that the evidence
is somewhat strained to justify this dogma.
In the first place, beyond the cases in which the nucleus, though
ordinarily invisible, is said to have been rendered visible by a re-agent,
there are cases, as in the already-named _Archerina_, where no re-agent
makes one visible. In the second place, there is the admitted fact that
some nuclei are diffused; as in _Trachelocerca_ and some other Infusoria.
In them the numerous scattered granules are supposed to constitute a
nucleus: an interpretation obviously biassed by the desire to save the
generalization. In the third place, the nucleus is frequently multiple in
cells of low types; as in some families of Algæ and predominantly among
Fungi. Once more, the so-called nucleus is occasionally a branching
structure scarcely to be called a "kernel."
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