The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
There are two kinds of homogenesis, the simplest of them, probably once
universal but now exceptional, being that in which there is no other form
of multiplication than one resulting from perpetual spontaneous fission.
The rise of distinct sexes was doubtless a step in evolution, and before it
took place the formation of new individuals could have arisen only by
division of the old, either into two or into many. At present this process
survives, so far as appears, among _Bacteria_, certain _Algæ_, and sundry
_Protozoa_; though it is possible that a rarely-occurring conjugation has
in these cases not yet been observed. It is a probable conclusion, however,
that in the _Bacteria_ at any rate, the once universal mode of
multiplication still survives as an exceptional mode. But now passing over
these cases, we have to note that the kind of genesis (once supposed to be
the sole kind), in which the successive generations are alike, is sexual
genesis, or, as it has been otherwise called--_gamogenesis_. In every
species which multiplies by this kind of homogenesis, each generation
consists of males and females; and from the fertilized germs they produce
the next generation of similar males and females arises: the only needful
qualification of this statement being that in many _Protophyta_ and
_Protozoa_ the conjugating cells or protoplasts are not distinguishable in
character. This mode of propagation has the further trait, that each
fertilized germ usually gives rise to but one individual--the product of
development is organized round one axis and not round several axes,
Homogenesis in contrast with heterogenesis as exhibited in species which
display distinct sexuality, has also the characteristic that each new
individual begins as an egg detached from the maternal tissues, instead of
being a portion of protoplasm continuous with them, and that its
development proceeds independently. This development may be carried on
either internally or externally; whence results the division into the
oviparous and the viviparous. The oviparous kind is that in which the
fertilized germ is extruded from the parent before it has undergone any
considerable development. The viviparous kind is that in which development
is considerably advanced, or almost completed, before extrusion takes
place. This distinction is, however, not a sharply-defined one: there are
transitions between the oviparous and the viviparous processes. In
ovo-viviparous genesis there is an internal incubation; and though the
young are in this case finally extruded from the parent in the shape of
eggs, they do not leave the parent's body until after they have assumed
something like the parental form. Looking around, we find that homogenesis
is universal among the _Vertebrata_. Every vertebrate animal arises from a
fertilized germ, and unites into its single individuality the whole product
of this fertilized germ. In the mammals or highest _Vertebrata_, this
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