The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
homogenesis is in every case viviparous; in birds it is uniformly
oviparous; and in reptiles and fishes it is always essentially oviparous,
though there are cases of the kind above referred to, in which viviparity
is simulated. Passing to the _Invertebrata_, we find oviparous homogenesis
universal among the _Arachnida_ (except the Scorpions, which are
ovo-viviparous); universal among the higher _Crustacea_, but not among the
lower; extremely general, though not universal, among Insects; and
universal among the higher _Mollusca_ though not among the lower. Along
with extreme inferiority among animals, we find homogenesis to be the
exception rather than the rule; and in the vegetal kingdom there appear to
be no cases, except among the _Algæ_ and a few aberrant parasites like the
_Rafflesiaceæ_, in which the centre or axis which arises from a fertilized
germ becomes the immediate producer of fertilized germs.
In propagation characterized by unlikeness of the successive generations,
there is asexual genesis with occasionally-recurring sexual genesis; in
other words--_agamogenesis_ interrupted more or less frequently by
_gamogenesis_. If we set out with a generation of perfect males and
females, then, from their ova arise individuals which are neither males nor
females, but which produce the next generation from buds. By this method of
multiplication many individuals originate from a single fertilized germ.
The product of development is organized round more than one centre or axis.
The simplest form of heterogenesis is that seen in most uniaxial plants.
If, as we find ourselves obliged to do, we regard each separate shoot or
axis of growth as a distinct individual, homogenesis is seen in those which
have absolutely terminal flowers; but in all other uniaxial plants, the
successive individuals are not represented by the series A, A, A, A, &c.,
but they are represented by the series A, B, A, B, A, B, &c. For in the
majority of plants which were classed as uniaxial (§ 50), and which may be
conveniently so distinguished from other plants, the axis which shoots up
from the seed, and substantially constitutes the plant, does not itself
flower but gives lateral origin to flowering axes. Though in ordinary
uniaxial plants the fructifying apparatus _appears_ to be at the end of the
primary, vertical axis; yet dissection shows that, morphologically
considered, each fructifying axis is an offspring from the primary axis.
There arises from the seed a sexless individual, from which spring by
gemmation individuals having reproductive organs; and from these there
result fertilized germs or seeds that give rise to sexless individuals.
That is to say, gamogenesis and agamogenesis alternate: the peculiarity
being that the sexual individuals arise from the sexless ones by continuous
development. The _Salpæ_ show us an allied form of heterogenesis in the
animal kingdom. Individuals developed from fertilized ova, instead of
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