The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Much more conspicuous are these general and special relations of form
to general and special actions in the environment, among phænogams
that are multiaxial. That when standing alone, and in places where
the winds do not injure them nor adjacent things shade them, shrubs
and trees develop with tolerable evenness on all sides, is an obvious
truth. Equally obvious is the truth that, when growing together in
a wood, and mutually interfered with on all sides, trees still show
obscurely radial distributions of parts; though, under such conditions,
they have tall taper stems with branches directed upwards--a difference
of shape clearly due to the different incidence of forces. And almost
equally obvious is the truth, that a tree of this same kind growing
at the edge of the wood, has its outer branches well developed and
its inner branches comparatively ill-developed. Fig. 197, which
inaccurately represents this difference, will serve to make it manifest
that while one of the peripheral trees can be cut into something like
two similar halves by a vertical plane directed towards the centre of
the wood--a plane on each side of which the conditions are alike--it
cannot be cut into similar halves by any other plane. A like divergence
from an indefinitely-radial symmetry towards an indefinitely-bilateral
symmetry, occurs in trees that have their conditions made bilateral
by growing on inclined surfaces. Two of the common forms observable
in such cases are given in Fig. 198. Here there is divisibility into
parts that are tolerably similar, by a vertical plane running directly
down the hill; but not by any other plane. Then, further, there is the
bilateralness, similar in general meaning though differently caused,
often seen in trees exposed to strong prevailing winds. Almost every
sea-coast has abundant examples of stunted trees which, like the one
shown in Fig. 199, have been made to deviate from their ordinary equal
growth on all sides of a vertical axis, to a growth that is equal only
on the opposite sides of a vertical plane directed towards the wind’s
eye.
From among vegetal aggregates of the third order, we have now only to
add examples of the entirely asymmetrical form which accompanies an
entirely irregular distribution of incident forces. Creeping plants
furnish such examples. They show, both when climbing up vertical or
inclined surfaces and when trailing on the ground, that their branches
grow hither and thither as the balance of forces aids or opposes;
and the general outline is without symmetry of any kind, because the
environing influences have no kind of regularity in their arrangement.
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