The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 220. Along with some unfamiliar facts, I have here set down facts
which are so familiar as to seem scarcely worth noting. It is because
these facts have become meaningless to perceptions deadened by infinite
repetitions of them, that it is needful here to point out their
meanings. Not alone for its intrinsic importance has the unlikeness
between the attached ends and the free ends been traced among plants
of all degrees of integration. Nor is it simply because of the
significance they have in themselves, that instances have been given
of those varieties of symmetry and asymmetry which the free ends of
plants equally display: be they plants of the first, second, third,
or any higher order. Neither has the only other purpose been that of
showing how, in the radial symmetry of some vegetal aggregates and
the single bilateral symmetry of others, there are traceable the same
ultimate principles as in the spherical symmetry and triple bilateral
symmetry of certain minute plants first described. But the main object
has been to present, under their simplest aspects, those general laws
of morphological differentiation which are fulfilled by the component
parts of each plant.
If organic form is determined by the distribution of forces, and the
approach in every case towards an equilibrium of inner actions with
outer actions; then this relation between forms and forces must hold
alike in the organism as a whole in its proximate units, and in its
units of lower orders. Formulas which express the shapes of entire
plants in terms of surrounding conditions, must be formulas which also
express the shapes of their several parts in terms of surrounding
conditions. If, therefore, we find that a plant as a whole is radially
symmetrical or bilaterally symmetrical or asymmetrical, according as
the incident forces affect it equally on all sides of an axis, or
affect it equally only on the opposite sides of one plane, or affect
it equally in no two directions; then, we may expect that, in like
manner, each member of a plant will display radial symmetry where
environing influences are alike along many radii, bilateral symmetry
where there is bilateralness of environing influences, and unsymmetry
or asymmetry where there is partial or entire departure from a balance
of surrounding actions.
To show that this expectation is borne out by the facts, will be the
object of the following four chapters. Let us begin with the largest
parts into which plants are divisible; and proceed to the successively
smaller parts.
CHAPTER VIII.
THE SHAPES OF BRANCHES.
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