The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 223. These few cases introduce us to the more familiar but more
complex cases which plants of the third degree of aggregation present.
At _a_, _b_, _c_, Fig. 202, are sketched three homologous parts of the
same tree: _a_ being the leading shoot; _b_ a lateral branch near
the top, and _c_ a lateral branch lower down. There is here a double
exemplification. While the branch _a_, as a whole, has its branchlets
arranged with tolerable regularity all round, in correspondence with
its equal exposure on all sides, each branchlet shows by its curve
as much bilateral symmetry as its simple form permits. The branch
_b_, dissimilarly circumstanced on the side next the main stem and
on the side away from it, has an approximate bilateralness as a
whole, while the bilateralness of its branchlets varies with their
respective positions. And in the branch _c_, having its parts still
more differently conditioned, these traits of structure are still more
marked. Extremely strong contrasts of this kind occur in trees having
very regular modes of growth. The uppermost branches of a Spruce-fir
have radially-arranged branchlets: each of them, if growing vigorously,
repeats the type of the leading shoot, as shown in Fig. 203, _a_,
_b_. But if we examine branches lower and lower down the tree, we
find the vertically-growing branchlets bear a less and less ratio to
the horizontally-growing ones; until, towards the bottom, the radial
arrangement has wholly merged into the bilateral. Shaded and confined
by the branches above them, these eldest branches develop their
offshoots in those directions where there is most space and light:
becoming finally quite flattened and fan-shaped, as shown at Fig.
203, _c_. And on remembering that each of these eldest branches, when
first it diverged from the main stem, was radial, we see not only that
between the upper and lower branches does this contrast in structure
hold, but also that each branch is transformed from the radial to
the bilateral by the progressive change in its environment. Other
forces besides those which aid or hinder growth, conspire to produce
this two-sided character in lateral branches. The annexed Fig. 204,
sketched from an example of the _Pinus Coulterii_ at Kew, shows very
clearly how, by mere gravitation, the once radially-arranged branchlets
may be so bent as to produce in the branch as a whole a decided
bilateralness. A full-grown _Araucaria_, too, exhibits in its lower
branches modifications similarly caused; and in each of such branches
there may be remarked the further fact, that its upward-bending
termination has a partially-modified radialness, at the same time that
its drooping lateral branchlets give to the part nearer the trunk a
completely bilateral character.
[Illustration: Figs. 202–204.]
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