The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Now in these few instances, typical of countless instances which might
be given, we see, as we saw in the case of the fungi, that the same
thing is true of the parts in their relations to the whole and to one
another, which is true of the whole in its relations to the environment
at large. Entire trees become bilateral instead of radial, when exposed
to forces that are equal only on opposite sides of one plane; and in
their branches, parallel changes of form occur under parallel changes
of conditions.
§ 224. There remains to be said something respecting the distribution
of leaves. How a branch carries its leaves constitutes one of
its characters as a branch, and is to be considered apart from
the characters of the leaves themselves. The principles hitherto
illustrated we shall here find illustrated still further.
The leading shoot and all the upper twigs of a fir-tree, have their
pin-shaped leaves evenly distributed all round, or placed radially;[32]
but as we descend we find them beginning to assume a bilateral
distribution; and on the lower, horizontally-growing branches, their
distribution is quite bilateral.[33] Between the Irish and English
kinds of Yew, there is a contrast of like significance. The branches
of the one, shooting up as they do almost vertically, are clothed with
leaves all round; while those of the other, which spread laterally,
bear their leaves on the two sides. In trees with better-developed
leaves, the same principle is more or less manifest in proportion as
the leaves are more or less enabled by their structures to maintain
fixed positions. Where the foot-stalks are long and slender, and where,
consequently, each leaf, according to its weight, the flexibility and
twist of its foot-stalk, and the direction of the branch it grows from,
falls into some indefinite attitude, the relations are obscured. But
where the foot-stalks are stiff, as in the Laurel, it will be found, as
before, that from the topmost and upward growing branches the leaves
diverge on all sides; while the undermost branches, growing out from
the shade of those above, have their leaves so turned as to bring them
into rows horizontally spread out on the two sides of each branch.
[Illustration: Fig. 205.]
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