The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Compound leaves that are completely bilateral, present us with
modifications of form exemplifying the same general truth in another
way. In them the proximal and distal parts have none of that
resemblance which we see in those intermediate forms just described.
The portion next the axis and the portion furthest from the axis
are entirely different; and the only likeness is between the wings
or leaflets on opposite sides of the main foot-stalk or mid-rib. On
turning back to Fig. 65, it will be seen that the compound leaf there
drawn to exemplify another truth, serves also to exemplify this truth:
the homologous parts _a_, _b_, _c_, _d_, while they are unlike one
another, are, in their main proportions, severally like the parts with
which they are paired. And here let us not overlook a characteristic
which is less conspicuous but not less significant. Each of the lateral
wings has winglets that are larger on the one side than on the other;
and in each case the two sides are dissimilarly conditioned. Even in
the several components of each wing may be traced a like divergence
from symmetry, along with a like inequality in the relations to the
rest: the proximal half of each leaflet is habitually larger than the
distal half. In the leaves of the Bramble, previously figured, kindred
facts are presented. How far such differences of development are due to
the positions of the parts in the bud; how far the respective spaces
available for the parts when unfolded affect them; and how far the
parts are rendered unlike by unlikenesses in their relations to light;
it is difficult to say. Probably these several factors operate in all
varieties of proportion. That the habitual shading of some parts by
others largely aids in causing these divergences from symmetry, is very
instructively shown by the compound leaves of the Cow-parsnip. Fig.
211 represents one of these. While the leaf as a whole is bilaterally
symmetrical, each of the wings has an unsymmetrical bilateralness: the
side next the axis being larger than the remoter side. How does this
happen? Fig. 212, which is a diagrammatic section down the mid-rib
of the leaf, showing its inclined attitude and the positions of the
wings _a_, _b_, _c_, will make the cause clear. As the wings overlap,
like the bars of a Venetian blind, each intercepts some light from the
one below it; and the one below it thus suffers more on its distal
side than on its proximal side. Hence the smaller development of the
distal side. That this is the cause is further shown by the proportion
that is maintained between the degree of obscuration and the degree
of non-development; for this unlikeness is greater between the two
sides _a_ and _a′_, than between _b_ and _b′_ or _c_ and _c′_, at the
same time that the interference is greater in the lower wings than in
the upper. Of course in this case and in the kindred cases hereafter
similarly interpreted, it is not meant that this differentiation
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