The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
is consequent solely, or even chiefly, on the differential actions
experienced by the individual plant. Though there is good reason to
believe that the rate of growth in each part of each leaf is affected
by the incidence of light, yet contrasts so marked and so systematic as
these are not explicable without taking into account the inheritance
of modifications either functionally caused or caused by spontaneous
variation. Clearly, the tendency will be towards the preservation of a
plant which distributes its chlorophyll in the most advantageous way;
and hence there will always be a gravitation towards a form in which
shaded parts of leaves are undeveloped.
§ 229. From compound leaves to simple ones, we find transitions in
leaves of which the divisions are partial instead of total; and in
these we see, with equal clearness, the relations between forms and
positions that have been traced thus far. Fig. 213 is the leaf of a
Winter-aconite in which, round a vertical petiole, there is a radial
distribution of half-separated leaflets. The _Cecropia_-leaf, Fig. 214,
shows us a two-sided development of the parts beginning to modify, but
not obliterating, the all-sided arrangement; and this mixed symmetry
occurs under conditions that are intermediate. A more marked degree of
the same relation is presented in the leaf of the Lady’s Mantle, Fig.
215. And then in the Sycamore and the Vine, we have a cleft type of
leaf in which a decided bilateralness of form co-exists with a decided
bilateralness of conditions.
[Illustration: Figs. 213–215.]
[Illustration: Figs. 216–218.]
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