The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 247. Keeping to the same type but passing to a higher degree of
composition, we meet more complex and varied illustrations of the same
general laws. In the compound _Cœlenterata_, presenting clusters
of individuals which are severally homologous with the solitary
individuals last dealt with, we have to note both the shapes of the
individuals thus united, and the shapes of the aggregates made up of
them.
[Illustration: Figs. 149, 150.]
Such of the fixed _Hydrozoa_ and _Actinozoa_ as form branched
societies, continue radial; both because their varied attitudes do not
expose them to appreciable differences in their relations to those
surrounding actions which chiefly concern them (the actions of prey),
and because such differences, even if they were appreciable, would
be so averaged in their effects on the dissimilarly-placed members
of each group as to be neutralized in the race. Among the tree-like
coral-polypedoms, as well as in such ramified assemblages of simpler
polypes as are shown in Figs. 149, 150, we have, indeed, cases in many
respects parallel to the cases of scattered flowers (§ 233), which
though placed laterally remain radial, because no differentiating
agency can act uniformly on all of them. Meanwhile, in the groups which
these united individuals compose, we see the shapes of plants further
simulated under a further parallelism of conditions. The attached ends
differ from the free ends as they do in plants; and the regular or
irregular branches obviously stand to environing actions in relations
analogous to those in which the branches of plants stand.
The members of those compound _Cœlenterata_ which move through the
water by their own actions, in attitudes that are approximately
constant, show us a more or less distinct two-sidedness. _Diphyes_,
Fig. 259, furnishes an example. Each of the largely-developed and
modified polypites forming its swimming sacs is bilateral, in
correspondence with the bilateralness of its conditions; and in each of
the appended polypites the insertion of the solitary tentacle produces
a kindred divergence from the primitive radial type. The aggregate,
too, which here very much subordinates its members, exhibits the same
conformity of structure to circumstances. It admits of symmetrical
bisection by a plane passing through its two contractile sacs, or
nectocalyces, but not by any other plane; and the plane which thus
symmetrically bisects it, is the vertical plane on the two sides of
which its parts are similarly conditioned as it propels itself through
the water.
[Illustration: Fig. 259.]
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