The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
the same part in advance--a form in which, consequently, the original
radial symmetry diverges more and more towards bilateral symmetry.
§ 250. Very definite and comparatively uniform, are the relations
between shapes and circumstances among the _Annulosa_: including under
that title the _Annelida_ and the _Arthropoda_. The agreements and the
disagreements are equally instructive.
At one time or other of its life, if not throughout its life, every
annulose animal is locomotive; and its temporary or permanent
locomotion, being carried on with one end habitually foremost and one
surface habitually uppermost, it fulfils those conditions under which
bilateral symmetry arises. Accordingly, bilateral symmetry is traceable
throughout the whole of this sub-kingdom. Traceable, we must say,
because, though it is extremely conspicuous in the immense majority
of annulose types, it is to a considerable extent obscured where
obscuration is to be expected. The embryos of the _Tubicolæ_, after
swimming about a while, settle down and build themselves tubes, from
which they protrude their heads; and in them, or in some of them, the
bilateral symmetry is disguised by the development of head-appendages
in an all-sided manner. The tentacles of _Terebella_ are distributed
much in the same way as those of a polype. The breathing organs in
_Sabella unispira_, Fig. 260, do not correspond on opposite sides of
a median plane. Even here, however, the body retains its primitive
bilateralness; and it is further to be remarked that this loss of
bilateralness in the external appendages, does not occur where the
relations to external conditions continue bilateral: witness the
_Serpula_, Fig. 261, which has its respiratory tufts arranged in a
two-sided way, under the two-sided conditions involved by the habitual
position of its tube.
[Illustration: Figs. 260–261.]
The community of symmetry among the higher _Annulosa_, has an
unobserved significance. That Flies, Beetles, Lobsters, Centipedes,
Spiders, Mites, have in common the characters, that the end which moves
in advance differs from the hinder end, that the upper surface differs
from the under surface, and that the two sides are alike, is a truth
received as a matter of course. After all that has been said above,
however, it will be seen to have a meaning not to be overlooked; since
it supplies a million-fold illustration of the laws which have been set
forth. It is needless to give diagrams. Every reader can call to mind
the unity indicated.
[Illustration: Figs. 263–270.]
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