The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
While, however, annulose animals repeat so uniformly these traits of
structure, there are certain other traits in which they are variously
contrasted; and their contrasts have to be here noted, as serving
further to build up the general argument. In them we see the stages
through which bilateral symmetry becomes gradually more marked, as
the conditions it responds to become more decided. A common Earth-worm
may be instanced as a member of this sub-kingdom that is among the
least-conspicuously bilateral. Though internally its parts have a
two-sided arrangement; and though the positions of its orifices give
it an external two-sidedness, at the same time that they establish
a difference between the two ends; yet its two-sidedness is not
strongly-marked. The form deviates but little from what we have
distinguished as triple bilateral symmetry: if the creature is cut
across the middle, the head and tail ends are very much alike; if
cut in two along its axis by a horizontal plane, the under and upper
halves are very much alike, externally if not internally; and if cut
in two along its axis by a vertical plane, the two sides are quite
alike. Figs. 263 and 264 will make this clear. Such creatures as
the _Julus_ and the Centipede, may be taken as showing a transition
to double bilateral symmetry. Besides being divisible into exactly
similar halves by a vertical plane passing through its axis, one of
these animals may be bisected transversely into parts that differ only
slightly; but if cut in two by a horizontal plane passing through its
axis, the under and upper halves are decidedly unlike. Figs. 265, 266,
exhibit these traits. Among the isopodous crustaceans, the departure
from these low types of symmetry is more marked. As shown in Figs. 267
and 268, the contrast between the upper and under parts is greater,
and the head and tail ends differ more obviously. In all the higher
_Arthropoda_, the unlikeness between the front half and the hind half
has become conspicuous. There is in them single bilateral symmetry of
so pronounced a kind, that no other resemblance is suggested than that
between the two sides. By Figs. 269 and 270, representing a decapodous
crustacean divided longitudinally and transversely, this truth is made
manifest. On calling to mind the habits of the creatures here drawn and
described, it will be seen that they explain these forms. The incidence
of forces is the same all around the Earth-worm as it burrows through
the compact ground. The Centipede, creeping amid loose soil or _débris_
or beneath stones, insinuates itself between solid surfaces--the
interstices being mostly greater in one dimension than in others.
And all the higher _Annulosa_, moving about as they do over exposed
objects, have their dorsal and ventral parts as dissimilarly acted upon
as are their two ends.
[Illustration: Fig. 271.]
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