The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Thus far, these aggregates of the second order exhibit but indefinite
individualities. The integration is physical; but not physiological.
Though, in the _Polycytharia_, there is a shape that has some symmetry;
and though, in the _Foraminifera_, the formation of successive chambers
proceeds in such methodic ways as to produce quite-regular and
tolerably-specific shells; yet no more in these than in the Sponges or
the compound _Vorticellæ_, do we find such co-ordination as gives the
whole a life predominating over the lives of its parts. We have not
yet reached an aggregate of the second order, so individuated as to be
capable of serving as a unit in still higher combinations. But in the
class _Cœlenterata_, this advance is displayed. The common _Hydra_,
habitually taken as the type of the lowest division of this class,
has specialized parts performing mutually-subservient functions, and
thus exhibiting a total life distinct from the lives of the units.
Fig. 146 represents one of these creatures in its contracted state
and in its expanded state; while Fig. 147 is a diagram showing the
wall of this creature’s sac-like body as seen in section under the
microscope: _a_ and _b_ being the outer and inner cellular layers;
while between them is the “mesoglœa” or “structureless lamella,” the
supporting or skeletal layer. But this lowly-organized tissue of the
Hydra, illustrates a phase of integration in which the lives of the
minor aggregates are only partially-subordinated to the life of the
major aggregate formed by them. For a _Hydra’s_ substance is separable
into _Amœba_-like portions, capable of moving about independently.
If we bear in mind how analogous are the extreme extensibility and
contractility of a _Hydra’s_ body and tentacles, to the properties
displayed by the sarcode among Rhizopods; we may infer that probably
the movements and other actions of a _Hydra_, are due to the
half-independent co-operation of the _Amœba_-like individuals composing
it.
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