The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Among the Ascidians this general law of morphological composition is
once more displayed. Each of these creatures subsists on the nutritive
particles contained in the water which it draws in through one orifice
and sends out through another; and it may thus subsist either alone, or
in connexion with others that are in some cases loosely aggregated and
in other cases closely aggregated. Fig. 156, _Phallusia mentula_, is
one of the solitary forms. A type in which the individuals are united
by a stolon that gives origin to them by successive buds, is shown in
_Perophora_, Fig. 157. Among the _Botryllidæ_, of which one kind is
drawn on a small scale in Fig. 159, and a portion of the same on a
larger scale in Fig. 158, there is a combination of the individuals
into annular clusters, which are themselves imbedded in a common
gelatinous matrix. And in this group there are integrations even a
stage higher, in which several such clusters of clusters grow from
a single base. Here the compounding and recompounding appears to be
carried further than anywhere else in the animal kingdom.
Thus far, however, among these aggregates of the third order, we see
what we before saw among the simpler aggregates of the second order--we
see that the component individualities are but to a very small extent
subordinated to the individuality made up of them. In nearly all the
forms indicated, the mutual dependence of the united animals is so
slight, that they are more fitly comparable to societies, of which
the members co-operate in securing certain common benefits. There is
scarcely any specialization of functions among them. Only in the last
type described do we see a number of individuals so completely combined
as to simulate a single individual. And even here, though there appears
to be an intimate community of nutrition, there is no physiological
integration beyond that implied in several mouths and stomachs having a
common vent.[20]
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