The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 204. We come now to an extremely interesting question. Does there
exist in other sub-kingdoms composition of the third degree, analogous
to that which we have found so prevalent among the _Cœlenterata_ and
the _Polyzoa_ and _Tunicata_? The question is not whether elsewhere
there are tertiary aggregates produced by the branching or clustering
of secondary aggregates, in ways like those above traced; but whether
elsewhere there are aggregates which, though otherwise unlike in the
arrangement of their parts, nevertheless consist of parts so similar to
one another that we may suspect them to be united secondary aggregates.
The various compound types above described, in which the united animals
maintain their individualities so distinctly that the individuality
of the aggregate remains vague, are constructed in such ways that the
united animals carry on their several activities with scarcely any
mutual hindrance. The members of a branched _Hydrozoon_, such as is
shown in Fig. 149 or Fig. 150, are so placed that they can all spread
their tentacles and catch their prey as well as though separately
attached to stones or weeds. Packed side by side on a flat surface or
forming a tree-like assemblage, the associated individuals among the
_Polyzoa_ are not unequally conditioned: or if one has some advantage
over another in a particular case, the mode of growth and the relations
to surrounding objects are so irregular as to prevent this advantage
re-appearing with constancy in successive generations. Similarly
with the Ascidians growing from a stolon or those forming an annular
cluster: each of them is as well placed as every other for drawing in
the currents of sea-water from which it selects its food. In these
cases the mode of aggregation does not expose the united individuals
to multiform circumstances; and therefore is not calculated to produce
among them any structural multiformity. For the same reason no marked
physiological division of labour arises among them; and consequently no
combination close enough to disguise their several individualities. But
under converse conditions we may expect converse results. If there is a
mode of integration which necessarily subjects the united individuals
to unlike sets of incident forces, and does this with complete
uniformity from generation to generation, it is to be inferred that
the united individuals will become unlike. They will severally assume
such different functions as their different positions enable them
respectively to carry on with the greatest advantage to the assemblage.
This heterogeneity of function arising among them, will be followed by
heterogeneity of structure; as also by that closer combination which
the better enables them to utilize one another’s functions. And hence,
while the originally-like individuals are rendered unlike, they will
have their homologies further obscured by their progressing fusion into
an aggregate individual of a higher order.
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