The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Very numerous are the heterogeneities of form presenting themselves
for interpretation under these heads. The ultimate morphological
units combined in any group, may be differentiated individually, or
collectively, or both: each of them may undergo changes of shape;
or some of them may be changed and others not; or the group may be
rendered multiform by the greater growth of some of its units than of
others. Similarly with the compound units arising by union of these
simple units. Aggregates of the second order may be made relatively
complex in form, by inequalities in the rates of multiplication of
their component units in diverse directions; and among a number of such
aggregates, numerous unlikenesses may be constituted by differences in
their degrees of growth, and by differences in their modes of growth.
Manifestly, at each higher stage of composition the possible sources of
divergence are multiplied still further.
That facts of this order can be accounted for in detail is not to
be expected--the data are wanting. All that we may hope to do is to
ascertain their general laws. How this is to be attempted we will now
consider.
§ 178. The task before us is to trace throughout these phenomena
the process of evolution; and to show how, as displayed in them, it
conforms to those first principles which evolution in general conforms
to. Two sets of factors have to be taken into account. Let us look at
them.
The factors of the first class are those which tend directly to
change an organic aggregate, in common with every other aggregate,
from that more simple form which is not in equilibrium with incident
forces, to that more complex form which is in equilibrium with them.
We have to mark how, in correspondence with the universal law that
the uniform lapses into the multiform, and the less multiform into
the more multiform, the parts of each organism are ever becoming
further differentiated; and we have to trace the varying relations
to incident forces by which further differentiations are entailed.
We have to observe, too, how each primary modification of structure,
induced by an altered distribution of forces, becomes a parent of
secondary modifications--how, through the necessary multiplication of
effects, change of form in one part brings about changes of form in
other parts. And then we have also to note the metamorphoses constantly
being induced by the process of segregation--by the gradual union
of like parts exposed to like forces, and the gradual separation of
like parts exposed to unlike forces. The factors of the second class
which we have to keep in view throughout our interpretations, are
the formative tendencies of organisms themselves--the proclivities
inherited by them from antecedent organisms, and which past processes
of evolution have bequeathed. We have seen it to be inferable from
various orders of facts (§§ 65, 84, 97–97_g_), that organisms are
built up of certain highly-complex molecules, which we distinguished as
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