The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
it finds nutriment, if its transverse section is flattened instead of
square or circular. And while we see how, by survival of the fittest, a
flattened form is likely to be acquired by diatoms having this habit;
we also see that likeness will be maintained between the two flat
surfaces and between the two edges. For, on the average, the relations
of the two flat surfaces to the sides of the openings through which
the diatom passes, will be alike; and so, too, on the average, will be
the relations of the two edges. In desmids of the type exemplified by
the second individual in Fig. 2, a kindred equalization of dimensions
is otherwise insured. There is nothing to keep one of the two surfaces
uppermost rather than the other; and hence, in the long succession
of individuals, the two surfaces are sure to be similarly exposed to
light and agencies in general. When to this is added the fact that
spontaneous fission occurs transversely in a constant way, it becomes
manifest that the two ends, while they are maintained in conditions
like one another, are maintained in conditions unlike those of the
two edges. Here then, as before, triple bilateral symmetry in form,
co-exists with a triple bilateral symmetry in the average distribution
of actions.
[Illustration: Figs. 4, 5, 6.]
Still confining our attention to aggregates of the first order, let us
next note what results when the two ends are permanently subject to
different conditions. The fixed unicellular plants, of which examples
are given in Figs. 4, 5, and 6, severally illustrate the contrast
in shape arising between the part that is applied to the supporting
surface and the part that extends into the surrounding medium. These
two parts which are the most unlike in their relations to incident
forces, are the most unlike in the forms. Observe, next, that the part
which lifts itself into the water or air, is more or less decidedly
radial. Each outward-growing tubule of _Codium adhærens_, Fig. 4, has
its parts disposed with some regularity around its axis; the upper stem
and spore-vessel of _Botrydium_, Fig. 5, display a lateral growth
that is approximately equal in every direction; and the stems of the
_Mucor_, Fig. 6, shoot up with an approach to evenness on all sides.
Plants of this low type are naturally very variable in their modes of
growth: each individual being greatly modified in form by its special
circumstances. But they nevertheless show us a general likeness between
parts exposed to like forces, as well as a general unlikeness between
parts exposed to unlike forces.
Respecting the forms of these aggregates of the first order, it has
only to be added that they are asymmetrical where there is total
irregularity in the incidence of forces. We have an example in the
indefinitely contorted and branched shape of a fungus-cell, growing as
a mycelium among the particles of soil or through the interstices of
organic tissue.
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