The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 218. Re-illustrations of the general truths which the forms of
these vegetal aggregates of the first order display, are furnished by
vegetal aggregates of the second order. The equalities and inequalities
of growth in different directions, prove to be similarly related to
the equalities and inequalities of environing actions in different
directions.
Of spherical symmetry an instance occurs in _Eudorina elegans_.
The ciliated cells are here so united as to produce a small,
mulberry-shaped, hollow ball which, being similarly conditioned on all
sides, shows no unlikenesses of structure. An allied form, however,
_Volvox globator_, presents a highly instructive, though very trifling,
modification. It is not absolutely homogeneous in its structure and
is not absolutely homogeneous in its motions. The waving cilia of
its component cells have fallen into such slight heterogeneities of
action as to cause rotation in a constant direction; and along with a
fixed axis of rotation there has arisen a fixed axis of progression. A
concomitant fact is that the cells of the colony exhibit an appreciable
differentiation in relation to the fixed axis. There is an incipient
divergence from spherical uniformity along with this slight divergence
from uniformity of conditions.
Vegetal aggregates of the second order are usually fixed: locomotion
is exceptional. Fixity implies that the surface of attachment is
differently circumstanced from the free surface. Hence we may expect
to find, as we do find, that among these rooted aggregates of the
second order, as among those of the first order, the primary contrast
of shape is between the adherent part and the loose part. Sea-weeds
variously exemplify this. In some the fronds are very irregular and
in some tolerably regular; in some the form is pseudo-foliar and in
some pseud-axial; but differing though they do in these respects,
they agree in having the end which is attached to a solid body unlike
the other end. The same truth is seen in such secondary aggregates
as the common Agarics, or rather in their immensely-developed organs
of fructification. A puff-ball, Fig. 192, presents no other obvious
unlikeness of parts than that between its under and upper surfaces. So
too with the stalked kinds that frequent our woods and pastures. In the
types which Figs. 193, 194, 195, delineate, the unlikenesses between
the rooted ends and the expanded ends, as well as between the under
and upper surfaces of the expanded ends, are obviously related to this
fundamental contrast of conditions. Nor is this relation less clearly
displayed in the sessile fungi which grow out from the sides of trees,
as shown at _a_, _b_, Fig. 196. That which is common to this and the
preceding types, is the contrast between the attached end and the free
end.
[Illustration: Figs. 192–196.]
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