Here we have a claim for the power of nature in the modification of species which fully comes up to the requirements of the most advanced evolutionist. It is remarkable, too, how clearly he perceived the great factors so important for the evolution of organisms, rapid multiplication, great variability, and the struggle for existence. Thus he remarks: “It may be said that the movement of nature turns upon two immovable pivots—one, the illimitable fecundity which she has given to all species; the other, the innumerable difficulties which reduce the results of that fecundity and leave throughout time nearly the same quantity of individuals in every species.” Here the term “difficulties” corresponds to the “positive checks” of Malthus, and to the “struggle for existence” of Darwin; and he again and again refers to variability—as when he says: “Hence, when by some chance, common enough with nature, a variation or special feature makes its appearance, man has tried to perpetuate it by uniting together the individuals in which it has appeared.” As Buffon thus clearly understood artificial selection, thoroughly appreciated the rapid increase of all organisms, and equally well saw that their inordinate increase was wholly neutralized through such destructive agencies as hunger, disease, and enemies, and as, at the same time, he had such unbounded faith in the power of nature to modify animal and vegetable forms, we feel assured that this great writer and original thinker only needed freedom to pursue this train of thought a little further and he would certainly have anticipated Darwin’s great discovery of natural selection by a whole century. Even as it is we must class him as one of the great pioneers of organic evolution. The next distinct step towards a theory of organic evolution was made by the poet Goethe at the very end of the eighteenth century, in his views of the metamorphosis of plants. He pointed out the successive modifications of the leaf which produced all the other essential parts of the higher plants—the simple cotyledons or seed leaves became modified into the variously formed leaves of the fully grown plants; these again were successively modified into the calyx, corolla, stamens, and ovary of the flower. He supposed this to be due to the increased refinement of the sap under the influence of light and air, and to indicate the steps by which the various parts of the flower had been developed. It was, therefore, a theory of evolution; but it was very unsatisfactory, inasmuch as it in no way accounted for the wonderful variety of the floral organs, or indicated any purpose served by the most prominent and conspicuous part of the flower, the highly colored and often strangely formed corolla. It was also erroneous in supposing that the corolla was a modified calyx, whereas it is now known to be a modification of the stamens.