The Dalmatian tribes and the Pannonians, at least of the region of the Save, for a short time obeyed the Roman governors; but they bore the new rule with an ever increasing grudge, above all on account of the taxes, to which they were unaccustomed, and which were relentlessly exacted. When Tiberius subsequently asked one of the leaders as to the grounds of the revolt, he answered that it had taken place because the Romans set not dogs and shepherds, but wolves, to guard their flocks. Now the legions from Dalmatia were brought to the Danube, and the men capable of arms were called out, in order to be sent thither to reinforce the armies. These troops made a beginning, and took up arms not for, but against, Rome. Their leader was one of the Daesitiatae (around Serajevo), Bato. The example was followed by the Pannonians, under the leadership of two Breuci, another Bato and Pinnes. All Illyricum rose with unheard of rapidity and unanimity. The number of the insurgent forces was estimated at 200,000 infantry and 9000 horsemen. The levy for the auxiliary troops, which had taken place more especially among the Pannonians to a considerable extent, had diffused more widely a knowledge of Roman warfare, along with the Roman language and even Roman culture. Those who had served as Roman soldiers formed now the nucleus of the insurrection.[14] The Roman citizens settled or sojourning in large number in the insurgent regions, the merchants, and above all, the soldiers, were everywhere seized and slain. The independent tribes, as well as those of the provinces, entered into the movement. The princes of the Thracians, entirely devoted to the Romans, certainly brought their considerable and brave bands to the aid of the Roman generals; but from the other bank of the Danube the Dacians, and with them the Sarmatae, broke into Moesia. The whole wide region of the Danube seemed to have conspired to put an abrupt end to the foreign rule.