One could easily imagine this mutilation to be the result of the
insect’s hard work. Poking about, digging and raking and slicing, at
one time in the gravelly soil, at another in the stringy mass of
manure, do not constitute a task in which organs so delicate as the
tarsi can be employed without risk. And here is an even more serious
matter: when the Beetle is rolling his ball backwards, with his head
down, it is with the extremities of his fore-feet that he presses
against the ground. What might not happen to the insect’s feeble
fingers, slender as thread, in consequence of this continual rubbing
against the rough soil? They are merely useless encumbrances; one day
or other they seem bound to disappear, crushed, torn off, worn out in a
thousand ways. We know unfortunately that our own workmen are all too
frequently injured in handling heavy tools and lifting great weights;
even so might the Scarab be crippled in rolling his ball, an enormous
load to him. In that case his maimed arms would be a noble testimony to
his industrious life.
But straightway grave doubts begin to assail us. If these mutilations
were really accidental and the result of too strenuous work, they would
be the exception, not the rule. Because a workman or several workmen
have had a hand caught and crushed in a machine, it does not follow
that all the rest will also lose their hands. If the Scarab sometimes,
or even very frequently, loses his fore-fingers in pursuing his trade
as a pill-roller, there must be some at least who, more fortunate or
more skilful, have preserved their tarsi. Let us then consult the
actual facts. I have observed in very large numbers the various species
of Scarabs that inhabit France: Scarabæus sacer, who is common in
Provence; S. semipunctatus, who keeps fairly close to the sea and
frequents the sandy shores of Cette, Palavas and the Golfe Juan;
lastly, S. laticollis, who is much more widely distributed than either
of the others and is found up the Rhone Valley at least as far as
Lyons. In addition, I have studied an African species, S. cicatricosus,
picked up near Constantine. Well, in all four species, the absence of
tarsi on the front legs has been an invariable fact, with not a single
exception, at any rate within the range of my observations. The Scarab
therefore is maimed from the start; and it is a natural peculiarity in
his case, not an accident.
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