Now all these difficulties vanish, if we consider the regeneration of
animals, such for instance as many worms of the annelid class or our
familiar ascidian *Clavellina*, in which regeneration in both directions
is possible. The wound at the posterior end of the one half which
results from the operation forms a posterior body half, the wound at
the anterior end of the other half forms an anterior one. Again, it is
the ideal whole which we meet here: each section of the body indeed may
be said to contain the potencies for the production of the totality,
though actually this totality is always realised by the addition of two
partial organisations. The title of complex-equipotential systems thus
seems to be fully justified as applied to the systems which are the
basis of regeneration: each section of the regenerating body may in fact
produce the same complex whole, or may, if we prefer to say so, at least
prepare the ground for that complex Anlage, out of which the complex
totality is actually to arise, in the same manner.
It often occurs in science, that in rather strange and abnormal
conditions something becomes apparent which might have been found
everywhere, which is lying before our eyes quite obviously. Are
we not in just such a condition at present? In order to study the
complex-equipotential systems, we turn to the phenomena of regeneration
and of restitution in general; we occasionally even introduce hypotheses
to render our materials more convenient for our purposes; and all the
time there is one sort of complex-equipotential system in the body of
every living being, which only needs to be mentioned in order to be
understood as such, and which indeed requires no kind of preliminary
discussion. The system of the propagation cells, in other words the
sexual organ, is the clearest type of a complex-equipotential system
which exists. Take the ovary of our sea-urchin for instance, and there
you have a morphogenetic system every element of which is equally
capable of performing the same complex morphogenetic course--the
production of the whole individual.
Further on we shall deal exclusively with this variety of our systems,
and in doing so we shall be brought back to our problem of heredity. But
it had its uses to place our concept of the complex-equipotential system
upon such a broad basis: we at once gave a large range of validity to
all that is to follow--which, indeed, does not apply to inheritance
alone, though its significance in a theory of heredity may be called its
most important consequence.
THE SECOND PROOF OF LIFE-AUTONOMY. ENTELECHY AT THE BOTTOM OF INHERITANCE
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