In denying any real explanatory value to the concept of natural
selection I am far from denying the action of natural selection. On
the contrary, natural selection, to some degree, is *self-evident*;
at least as far as it simply states that what is incompatible with
permanent existence cannot exist permanently, it being granted that
the originating of organic individuals is not in itself a guarantee of
permanency. Chemical compounds, indeed, which decompose very rapidly
under the conditions existing at the time when they originated may also
be said to have been eliminated by “natural selection.” It is another
question, of course, whether in fact all eliminations among organic
diversities are exclusively due to the action of natural selection in
the proper Darwinian sense. It has been pointed out already by several
critics of Darwinism and most clearly by Gustav Wolff, that there are
many cases in which an advantage with regard to situation will greatly
outweigh any advantage in organisation or physiology. In a railway
accident, for instance, the passengers that survive are not those who
have the strongest bones, but those who occupied the best seats; and
the eliminating effect of epidemics is determined at least as much by
localities, *e.g.* special houses or special streets, as by the degree
of immunity. But, certainly, natural selection is a *causa vera* in many
other cases.
We now may sum up our discussion of the first half of Darwinism.
Natural selection is a negative, an eliminating factor in transformism;
its action is self-evident to a very large degree, for it simply
states that things do not exist if their continuance under the given
conditions is impossible. To consider natural selection as a positive
factor in descent would be to confound the sufficient reason for the
non-existence of what is not, with the sufficient reason of what is.
Natural selection has a certain important logical bearing on
systematics, as a science of the future, which has scarcely ever been
alluded to. Systematics of course has to deal with the totality of the
possible, not only of the actual diversities; it therefore must remember
that more forms may be possible than are actual, the word “possible”
having reference in this connection to originating, not to surviving.
Moreover, systematics is concerned not only with what has been
eliminated by selection, but also with all that might have originated
from the eliminated types. By such reasoning natural selection gains a
very important aspect--but a logical aspect only.
FLUCTUATING VARIATION THE ALLEGED CAUSE OF ORGANIC DIVERSITY
Public-domain text, read in full here on John Shaqi.
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