There is finally one group of facts often brought forward against
Lamarckism by Darwinian authors[158] which may be called the logical
*experimentum crucis* of this doctrine, an *experimentum* destined
to prove fatal. You know that among the polymorphic groups of bees,
termites, and ants, there exists one type of individuals, or even
several types, endowed with some very typical features of organisation,
but at the same time absolutely excluded from reproduction: how could
those morphological types have originated on the plan allowed by the
Lamarckians? Of what use would “judgment” about means that are offered
by chance and happen to satisfy needs, be to individuals which die
without offspring? Here Lamarckism becomes a simple absurdity, just as
Darwinism resulted in absurdities elsewhere.
[158] It has also very often been said by Darwinians that Lamarckism is
only able to explain those cases of adaptedness which relate to active
functioning but not mere passive adapted characters, like “mimicry” for
example. But this argument *taken by itself*, it seems to me, would not
be fatal to Neo-Lamarckism in the special form August Pauly gave to this
doctrine.
We were speaking about dogmatic Darwinism then, and it is about dogmatic
Lamarckism that we are reasoning at present; both theories must fall in
their dogmatic form, though a small part of both can be said to stand
criticism. But these two parts which survive criticism, one offered by
Lamarck, the other by Darwin, are far from being a complete theory of
transformism, even if taken together: they only cover a small area of
the field concerned in the theory of descent. Almost everything is still
to be done, and we may here formulate, briefly at least, what we expect
to be accomplished by the science of the future.
4. THE REAL RESULTS AND THE UNSOLVED PROBLEMS OF TRANSFORMISM
What has been explained to a certain extent by the two great theories
now current is only this. Systematic diversities consisting in mere
differences as to intensity or number may perhaps owe their origin to
ordinary variation. They may at least, if we are entitled to assume
that heredity in some cases is able to hand on such variations without
reversion, which, it must be again remarked, is by no means proved by
the facts at present. Natural selection may share in this process by
eliminating all those individuals that do not show the character which
happens to be useful. That is the Darwinian part of an explanation of
transformism which may be conceded as an hypothesis. On the other side,
congenital histological adaptedness may be regarded hypothetically as
due to an inheritance of adaptive characters which had been acquired by
the organism’s activity, exerted during a great number of generations.
That is the Lamarckian part in the theory of descent.
Public-domain text, read in full here on John Shaqi.
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