But nothing more is contributed to this theory either by the doctrine
of Darwin or by that of Lamarck. So it follows that almost everything
has still to be done; for no hypothesis at present accounts for
the foundation of all systematics, viz., for the differences in
organisation, in all that relates to the so-called types as such and
the degree of complication in these types, both of which (types and
degree of complication) are independent of histological adaptation and
adaptedness.
What then do we know about any facts that might be said to bear on
this problem? We have stated already at the end of our chapter devoted
to the analysis of heredity that what we actually know about any
deviation of inheritance proper, that is, about congenital differences
between the parents and the offspring, relating to mere tectonics,
is practically nothing: indeed, there are at our disposal only the
few facts observed by de Vries or derived from the experience of
horticulturalists and breeders. We may admit that these facts at
least prove the possibility[159] of a discontinuous variation, that
is of “mutation,” following certain lines of tectonics and leading to
*constant* results; but everything else, that is everything about a real
theory of phylogeny, must be left to the taste of each author who writes
on the theory of the Living. You may call that a very unscientific state
of affairs, but no other is possible.
[159] But nothing more. All “mutations” hitherto observed in nature
or (comp. page 238, note 3) experimentally produced relate only to
“varieties” and not to “species.” One could hardly say that the
recent investigations about the production of mutations by *external*
means have strengthened their importance for the general theory of
transformism.
And, in fact, it has been admitted by almost all who have dealt with
transformism without prepossessions that such is the state of affairs.
Lamarck himself, as we have mentioned already, was not blind to the fact
that a sort of organisatory law must be at the base of all transformism,
and it is well known that hypothetical statements about an original law
of phylogeny have been attempted by Nägeli, Kölliker, Wigand, Eimer,
and many others. But a full discussion of all these “laws” would hardly
help us much in our theoretical endeavour, as all of them, it must be
confessed, do little more than state the mere fact that some unknown
principle of organisation must have been at work in phylogeny, if we are
to accept the theory of descent at all.
It is important to notice that even such a convinced Darwinian as
Wallace, who is well known to have been an independent discoverer of the
elimination principle, admitted an exception to this principle in at
least one case--with regard to the origin of man. But one exception of
course destroys the generality of a principle.
Public-domain text, read in full here on John Shaqi.
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