a lower side before: it now has got an upper and lower, front and back,
*right and left* half; it now has acquired that symmetry of organisation
which our own body has; at least it has got it as far as its mesenchyme
is concerned.
[Illustration: Fig. 3.--Formation of Mesenchyme in Echinus.
*a.* Outlines of blastula, side-view; mesenchyme forms a heap of cells
at the “vegetative” pole.
*a*_1. Heap of mesenchyme-cells from above.
*b.* Mesenchyme-cells arranged in a ring round the vegetative pole.
*c.* Mesenchyme-cells arranged in a bilateral-symmetrical figure;
primordia of skeleton in the midst of two spherical triangles.]
We leave the mesenchyme for a while and study another kind of
organogenesis. At the very same pole of the germ where the mesenchyme
cells originated there is a long and narrow tube of cell growing in,
and this tube, getting longer and longer, after a few hours of growth
touches the opposite pole of the larva. The growth of this cellular tube
marks the beginning of the formation of the intestine, with all that
is to be derived from it. The larva now is no longer a blastula, but
receives the name of “gastrula” in Haeckel’s terminology; it is built
up of the three “germ-layers” in this stage. The remaining part of the
blastoderm is called “ectoderm,” or outer layer; the newly-formed tube,
“endoderm,” or inner layer; while the third layer is the “mesenchyme”
already known to us.
The endoderm itself is a radial structure at first, as was the whole
germ in a former stage, but soon its free end bends and moves against
one of the sides of the ectoderm, against that side of it where the two
triangles of the mesenchyme are to be found also. Thus the endoderm
has acquired bilateral symmetry just as the mesenchyme before, and
as in this stage the ectoderm also assumes a bilateral symmetry
in its form, corresponding with the symmetrical relations in the
endoderm and the mesenchyme, we now may call the whole of our larva a
bilateral-symmetrical organisation.
It cannot be our task to follow all the points of organogenesis of
Echinus in detail. It must suffice to state briefly that ere long a
second portion of the mesenchyme is formed in the larva, starting from
the free end of its intestine tube; that the formation of the so-called
“coelum” occurs by a sort of splitting off from this same original
organ; and that the intestine itself is divided into three parts of
different size and aspect by two circular sections.
Public-domain text, read in full here on John Shaqi.
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