In Egil, as in Highland Chieftain, there
is also evidence of arrested growth in the facial part of the skull;
the profile is concave and the frontal index high (65) as in Highland
Chieftain.
Having seen that, apart from the face, the only essential difference
between the skeleton of an unimproved Shetland and the skeleton of an
Exmoor pony is a difference in size, let us next direct attention to
the skeleton of Eric, a 36·5-inch improved pedigree pony of the riding
type, for some time in the possession of Mr Charles M. Douglas of
Auchlochan. In Eric, who died when six years old, the fore limb from
the elbow to the ground was 22 inches, the length from the point of the
hock to the ground 15·4 inches, the circumference below the knee 5·25
inches, and the width of the fore-shank 1·2 inches. All four ergots
were present, but the hind chestnuts were absent. Though the setting-on
of the tail, the somewhat rounded hindquarters and the presence of
ergots, indicated that Eric, like practically all modern Shelties,
included horses of the “forest” type amongst his ancestors, his skull,
teeth, and limbs made it evident that he mainly belonged to the Celtic
or riding type.
In Eric the face (fig. 6) is so short that the frontal index is 67, the
length being only 1·4 times the width instead of 1·8 as in 12-hands
ponies of the riding type. The length of Eric’s head when alive was 410
mm. (16¼ inches). A typical Celtic pony with a 410 mm. head measures
40 inches at the withers. Eric, though having the head of at least a
40-inch pony, only measured 36·5 inches. It may hence be assumed that,
through dwarfing, his total height was reduced by 3·5 inches.
Further, as Eric’s metacarpal (fig. 9_a_), instead of measuring 166 mm.
(the normal length in a 40-inch pony), had only a length of 143 mm.,
it follows that practically one inch of the dwarfing was due to a
reduction in the length of the cannon-bones. Moreover, as Eric actually
measured 36·5 inches, his metacarpals should have measured 152 mm.
instead of 143 mm.—143 mm. being the length in a normal pony measuring
34 inches at the withers.
Though Eric had the head and trunk of a 40-inch pony and the
metacarpals of a 34-inch pony, the metacarpals bear the same relation
to the radius and the humerus as in Persimmon—_i.e._, in Eric the
relative lengths of the different parts of the limb were maintained
(not lost, as in the dachshund) during the dwarfing process.
Although the cannon-bones in Eric had been considerably reduced in
length, they had not been reduced in width—_i.e._, they are as wide as
the metacarpals of the 40-inch Shetland pony Egil, in which the limbs
closely conform to the Celtic type. It thus appears that, in the case
of ponies, reduction in height at the withers, and especially in the
length below the knee, is not necessarily accompanied by loss of “bone.”
Public-domain text, read in full here on John Shaqi.
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