Leaving the Typhlopidæ and Glauconiidæ aside for the present, snakes
have a wide gape, cleft far beyond the vertical of the eyes, with, when
closed, one or two notches in front for the passage of the protrusible,
bifid tongue. In most snakes this chink is in the lower border of the
rostral shield, capping the tip of the snout, and allows free passage to
the whole tongue; in the Hydrophids, or Sea-snakes, there are two
notches in the lower border of the rostral shield, through which only
the bifid end of the tongue can be protruded. The eyes, varying from
minute to enormous, are usually free from the surrounding shields, and
may move under a transparent cap like a watch-glass, which appears to
represent the lower eyelid of Lizards. The view as to this homology is
derived from our knowledge of various conditions in certain series of
Lizards of the families Lacertidæ and Scincidæ, where we find a
transparent disc appearing like a small window in the movable lower
eyelid, gradually increasing in size so as to occupy the whole of the
lower eyelid, which finally becomes fused with the rudimentary upper lid
and loses its mobility. In _Ilysia_ and in most of the Uropeltidæ, the
transparent disc over the eye is confluent with a thick horny shield of
which it occupies the middle.
The pupil is usually circular or vertical, rarely horizontal. In some
forms it is difficult to decide whether it is round or vertically
elliptic; in others, like the Boas and Vipers, for instance, it is
decidedly vertical, and contracts to the same extent as a cat's. In some
Water-snakes, and in Sea-snakes generally, the round pupil may contract
to a mere dot. The contraction of the pupil is independent on the two
sides.
The snout, or the part of the head anterior to the eyes, may be short or
long, rounded or pointed, depressed or compressed, sometimes projecting
strongly beyond the mouth, turned up at the end, or terminating in one
(_Langaha_) or two (_Herpeton_) long scaly dermal appendages. In some
burrowing forms it is provided with a more or less trenchant horizontal
or vertical edge. When the sides of the snout (loreal region) form an
angle with the upper surface, the angle is termed the "canthus
rostralis," which may be intensified by the loreal region being concave.
The deep pits which are sometimes present on the lips or between the
nostril and the eye (loreal pit) will be alluded to further on under
Sensory Organs.
The nostrils are either lateral, or, in the aquatic forms, directed
upwards, sometimes entirely on the upper surface of the snout.
Most snakes have a longitudinal groove on the chin (mental groove) to
allow for the distension caused by the lateral movements of the rami of
the lower jaw.
Public-domain text, read in full here on John Shaqi.
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