_cs_, Chin-shields (anterior); _cs´_, chin-shields (posterior); _f_,
frontal; _in_, internasal; _l_, loreal; _la_, labial (upper); _la´_,
labial (lower); _m_, mental; _n_, nasal; _p_, parietal; _pf_,
prefrontal; _pro_, preocular; _pto_, postocular; _r_, rostral;
_sbo_, subocular; _so_, supraocular; _t_, temporals (first row);
_t´_, temporals (second row); _v_, first ventral.]
The periodical shedding of the outer layer of the epidermis in a single
piece, including even the covering of the eye, is one of the most
striking peculiarities of snakes, although paralleled in the Lizards of
the family Anguidæ, to which our British Slow-worm belongs. The skin
becomes detached at the lips, and is turned inside out from head to
tail, without any sort of laceration when the snake is in good health.
These exuviæ are transparent, but often carry a certain amount of
pigment, especially those of the Vipers, in which the characteristic
dark markings are perfectly visible; they usually exceed the length of
the reptile, owing to stretching. In Sea-snakes the epidermis is cast
piecemeal, and sloughing is a longer operation than in ordinary snakes.
In Rattlesnakes each piece of the rattle, or "crotalon," in which the
tail terminates, represents a retained portion of the sloughed
epidermis. This remarkable appendage looks like a number of horny rings,
but it consists in reality of hollow, bell-like pieces, similar to the
terminal one, or "button," each with a circular constriction, in which
the incurved free edge of the following piece fits, thus keeping the
pieces together without impairing the mobility necessary to produce the
rattling sound for which the apparatus is intended. At each exuviation
one bell-shaped horny piece is added. The number of segments in the
rattle is, therefore, not an index to age, as formerly believed; nor is
it to the number of exuviations, for whilst segments are being added at
the base of the apparatus the terminal ones break off and are lost. A
_Crotalus_ sixteen months old may have six pieces to the rattle if there
have been six exuviations and no loss. No rattle appears ever to
comprise more than about twenty pieces, even in old specimens. The size
of the terminal button shows whether it was formed at birth or at any
later period, no growth taking place in the horny tissue.
Public-domain text, read in full here on John Shaqi.
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