The poison fangs of the Viperidæ appear to be movable, folding in the
mouth when at rest, and erected, or even thrust forward, when ready to
act. This, however, is simply due to the mobility of the maxillary bone,
to which they are ankylosed as in all other snakes. There are normally
two equally-developed fangs, close together and side by side, to each
maxillary, followed by several replacement fangs loosely attached behind
them, usually in two series of four. When the two fangs are _in situ_,
they of course both function in the act of biting, although only one is
in relation with the single poison duct; often, however, there is only
one fang in position, either the right or the left, the place of the
other being indicated by a shallow socket which will soon be filled by
one of the posterior reserve fangs moving forward and becoming ankylosed
to the bone. Snake-charmers who extract the poison fangs of the snakes
they use for their performances have therefore to renew the operation
frequently, unless they amputate the bone on which the fangs are
inserted, an injury which the creature does not long survive.
The dentition of the snakes in which the maxillary bone is not movable
vertically falls under three divisions: the Aglyphs, in which the teeth
are all solid; the Opisthoglyphs, in which one or more (usually two) of
the hindermost teeth are provided with a groove; and the Proteroglyphs,
in which grooved or canaliculated teeth are situated in front, followed
or not by solid teeth. Beyond these three principal divisions, the
dentition furnishes important characters for the classification,
although that importance has sometimes been over-estimated. The
maxillary teeth may be equal in length (Isodonts), or the anterior the
longer (Lycodonts), or the posterior the longer, increasing gradually in
size (Coryphodonts) or abruptly, without (Syncranterians) or with a
diastema, or break, in front of them (Diacranterians). These categories
are, however, so completely connected as to preclude their use in
taxonomy beyond helping to define genera. The number of maxillary teeth
and the relative proportions and disposition of the mandibular teeth
also afford useful generic characters.
The European genera may be arranged as follows, according to the
dentition:
I. Teeth few, disposed in a transverse series in the upper jaw only
_Typhlops._
II. Teeth in both jaws and on the palatines and pterygoids.
_A._ A series of solid teeth along the maxillary; no grooved teeth.
1. Anterior maxillary and mandibular teeth longest; 9 or 10
maxillary teeth
_Eryx._
2. Maxillary teeth equal, or increasing in size posteriorly.
_a._ Mandibular teeth 17 to 30; maxillary teeth 15 to 22.
Posterior maxillary teeth longest; mandibular teeth subequal, more than
20
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